Prosopocoilus (Metopodontus) WESTWOOD in HOPE, 1845

(G. = Lucanus downesii)


Prosopocoilus (Metopodontus) astacoides (HOPE, 1840).

Hope, 1840:78; Hope, 1841:299; Hope, 1841:590-591; Hope & Westwood, 1845:12; Waterhouse, 1869:17; Parry, 1870:78, pl.3; Boileau, 1911:63; Boileau, 1913:225; Gravely, 1915:418-419; Didier, 1927:158; Didier, 1928:70; Didier, 1929:121; Arrow, 1950:165, pl. 15; Didier & Séguy, 1952:pl.79; Didier & Séguy, 1953:41, 103, 105, 107; Benesh, 1960:63, 68; Endrodi, 1968:51; Kurosawa, 1970:161, pl.2; Nakane, 1972:42; De Lisle, 1975:204; Maes, 1981:3-4; Sakaino, 1981:23, 25; Biswas & Chatterjee, 1985:182; Kawano, 1989:19; Maes, 1990:6, 9; Bomans, 1991:146; Maes, 1992:69-70; Kawano, 1992:12; Kawano, 1993:12; Bartolozzi & Sforzi, 1994:109, 114; Mizunuma & Nagai, 1994:25-26, 28, 252, 253-254, pl. 71, 73-74; Kawano, 1997:454.

 

Prosopocoilus (Metopodontus) astacoides ssp. poultoni (BOILEAU, 1911).

Metopodontus poultoni BOILEAU, 1911 [Sakiou, Maria Basti].

Metopodontus croceus DIDIER 1929 [Bhutan].

Prosopocoilus astacoides ssp. astacoides (HOPE); Mizunuma & Nagai, 1994.

India : Uttar Pradesh (Almora--Kimoli), West Bengal (Darjeeling, Gopaldhara, Rungbong Valley, Mangpu, Kurseong, Pashok, Abor Country--Kobo), Assam;

Nepal : Likhu Khola-Tal, Junbesi, Jubing, Jiri, Balaju--Katmandu, Katmandu, Village Bodha, Taplejung Distr.--Mechi Zone--Dobhan;

Bhutan : Pedong, Sakiong = Sakiou*, Kaggia, Maria Basti*, Wangdi-Phodrang, Phodrang;

Thailand : Chiang Mai (Chom Thong--Maeklang, Mt. Doi Suthep).

 

Prosopocoilus (Metopodontus) astacoides ssp. astacoides (HOPE, 1840).

Lucanus astacoides HOPE, 1840 [Assam].

Lucanus foveatus HOPE, 1840 [Assam].

Lucanus omissus HOPE, 1840 [Assam].

Lucanus fraternus HOPE & WESTWOOD, 1845 [Loc. ?].

Cladognathus impressus WATERHOUSE, 1869 [India].

Prosopocoilus astacoides ssp. castaneus HOPE & WESTWOOD; Mizunuma & Nagai, 1994.

Prosopocoilus castaneus HOPE & WESTWOOD; Kawano, 1997.

India : Arunachal Pradesh (Namdapha, Miao), Assam*, Meghalaya (Khasi Hills, Cherrapunji, Shillong), Nagaland (Kohima, Naga Hills), Manipur;

Bangladesh : Sylhet;

Myanmar = Burma : Shan State (Keng Tung), ? (Thandaung);

Thailand : Chiang Rai (Wiangpapao), Chiang Mai (Chiang Mai--18°50N-98°57E, Mt. Doi Pui, Mt. Doi Inthanon--18°26N-98°17E, Doi Suthep), Meo--N. Thailand.

 

Prosopocoilus (Metopodontus) astacoides ssp. birmanicus (GRAVELY, 1915).

Metopodontus astacoides ssp. birmanicus (GRAVELY, 1915) [Haka, Rangoon].

Myanmar = Burma : Chin State (Chin Hills--Haka*), Shan States (Taunggyi--20°55N-96°49E); Rangoon State (Rangoon*--16°48N-96°08E), ? (Sadon);

Andaman Is.

 

Prosopocoilus (Metopodontus) astacoides ssp. kachinensis BOMANS & MIYASHITA, 1997.

Prosopocoilus astacoides ssp. kachinensis BOMANS & MIYASHITA, 1997:2, figs. 1, 2 [Kachin Prov.].

Myanmar :

Kachin Prov., VIII-96, leg. P. Sukkit (HOLOTYPUS in coll. Miyashita, Japan) (5 male PARATYPUS in coll. Miyashita, Japan; Bomans, in Natural History Museum, London; Arnaud, Francia).

Nota :

Bomans & Miyashita give that this subspecies differs from others by two firsts teeths very near each others and epistome with two small tubercules.

Psosopocoilus (Metopodontus) astacoides ssp. kachinensis (adapted from Bomans & Miyashita, 1997).

 

Bibliography :

BOMAN H. & MIYASHITA T. (1997) Description de trois nouvelles espèces de Lucanidae de nord Birmanie. 91 ème contribution à l'étude des Coléoptères Lucanides. Besoiro, 4:2-3.

 

 

Prosopocoilus (Metopodontus) bison (OLIVIER, 1789).

Herbst, 1790:298, pl.33; Fabricius, 1792:238, pl.1; Olivier (ed. Illiger), 1800:61; Fabricius, 1801:250; Olivier (ed. Sturm), 1802:2, pl.2; Thunberg, 1806:193; Voet, 1806:51, pl.30; Olivier, 1808:pl.3; Schonherr, 1817:324; Burmeister, 1847:373; Blanchard, 1853:138, pl.9; Montrouzier, 1857:27; Parry, 1864:22; Parry, 1870:108; Van Roon, 1905:115-116; Mollenkamp, 1906:31; Heyne & Taschenberg, 1908:53, pl.7; Van Roon, 1910:22-23; Boileau, 1913:225; Arrow, 1915:502; Kriesche, 1920:93; Kriesche, 1924:439-440; Nagel, 1930:91; Arrow, 1943:138; Didier & Séguy, 1952:pl.70; Didier & Séguy, 1953:40, 104-106; Benesh, 1960:64; De Lisle, 1964:8; Damoiseau, 1966:6; De Lisle, 1967:526; Bomans, 1971:334-351; Weinreich, 1971:226, pl.4; Weinreich, 1972; Maes, 1981:3; Klausnitzer, 1982:23; Lacroix, 1982:21; Bomans, 1984:354; Lacroix, 1988:12; Sakaino, 1988:17; Maes, 1990:6; Maes, 1992:70; Moore & Cassis, 1992:15; Haitlinger, 1993:342; Haitlinger, 1994:193; Suzuki, 1994:21-23; Mizunuma & Nagai, 1994:244-245, pl. 60; Bomans & Bartolozzi, 1996:220; Bartolozzi, 1996:132

Note : Olivier (1789) refers the species bison to Voet, Coleopt. tab. 30. fig. 6.

 

Prosopocoilus (Metopodontus) bison ssp. flavocinctus (WEINREICH, 1971) [Prosopocoelus].

Sumatra* : N. Sumatra Prov. (Doloc Baros Estate--Medan);

Java;

Sulawesi = Celebes.

 

Prosopocoilus (Metopodontus) bison ssp. buruensis (KRIESCHE, 1920) [Metopodontus].

Moluccas Is. : Buru I.*--3°22S-126°35E.

 

Prosopocoilus (Metopodontus) bison ssp. tesserarius (HERBST, 1790) [Lucanus, Metopodontus, Prosopocoelus, Metapodonotus] [Loc. ?].

= Lucanus fulvolimbatus BLANCHARD 1853 [Amboine].

Moluccas Is. : Ambon I.--3°39S-128°07E = Amboina I., Seram I.--3°05S-129°20E = Ceram I. (Pirae), Ternate I.--0°50N-127°15E, Banda I.;

Lesser Sunda Is. : Kai I.--5°42S-132°55E = Key I.;

New Guinea.

Natural enemies : Acarina : Canestriniidae : Amboinophela bernae.

 

Prosopocoilus (Metopodontus) bison ssp. magnificus (MOLLENKAMP, 1906) [Metopodontus] [Australia, error].

Lesser Sunda Is. : Tanimbar I.--7°31S-131°30E.

 

Prosopocoilus (Metopodontus) bison ssp. bison (OLIVIER, 1789).

Lucanus bison OLIVIER, 1789:13, pl.3 [America Meridionalis, error].

Lucanus cinctus MONTROUZIER, 1857 [Cladognathus, Metopodontus, Prosopocoelus] [Woodlarck I.].

Metopodontus tumidicollis MOLLENKAMP, 1906 [Prosopocoilus] [aumidicollis] [New Guinea : Astrolabe Bay].

Zacher, 1913.

Metopodontus monodon KRIESCHE, 1924 [Herbertshöhe].

Prosopocoilus (Metopodontus) bison (OLIVIER); Cammaerts & Bomans, 1997:13.

Lesser Sunda Is. : Kai I., 5°42S-132°55E

Lesser Sunda Is. : Aru I., 5°59S-134°30E

New Guinea : Irian Jaya : Mt. Arfak

New Guinea : Irian Jaya : Ransiki near Mt. Arfak

New Guinea : Irian Jaya : Manokwari Prov. : Testega

New Guinea : Irian Jaya : Mapia near Nabire

New Guinea : Irian Jaya : Numfor I., 1°00S-134°51E

New Guinea : Irian Jaya : Biak I.-- 0°53S-135°55E

New Guinea : Papua : Wau, 7°19S-146°37E

New Guinea : Papua : Wareo

New Guinea : Papua : Fly River

New Guinea : Papua : Sattelberg

New Guinea : Papua : Mt. Hagen : Baiyer River

New Guinea : Papua : Kielo : Arowa Plantation

New Guinea : Papua : Madang Province : Manam I. : Budua, alt. 20 m, 10-XII-1989, in coastal forest very damaged, on volcanic soil (1 male in col. H.E. Bomans, in Natural History Museum, London).

New Guinea : Papua, leg. Hariyanto (3 males, 1 female in col. Ronald Cave, USA).

New Guinea : Mt. Missim

New Guinea : Astrolabe Bay

New Guinea : Wamoro Mts.

New Guinea : Berlinhafen

New Guinea : Regenberg

New Guinea : Pulo Fahor

New Guinea : Soron

New Guinea : Mansinam

New Guinea : Hatam

New Guinea : Foce del Katau

New Guinea : Einsiedler I.

Bismarck Is. : Mussau I., 1°26S-149°36E : Talumalaus

Bismarck Is. : Mussau I., 1°26S-149°36E : Malakata, 1°28 S – 149°33 E

New Ireland I., 3°19S-151°55E : Lemkamin, 3°20 S – 151°55 E

New Britain I., 5°44S-150°41E = New Pommern I. : Gaz Halbins : Erbertshöhe

New Britain I., 5°44S-150°41E = New Pommern I. : Yalom, 4°25 S – 151°45 E.

Yule I.

D'Entrecasteaux Is. : Woodlarck I., 9°02S-152°43E.

Xylophagous : Arecaceae : Cocos.

Natural enemies : Acarina : Canestriniidae : Amboinophela bernae.

Bibliography.

CAMMAERTS R. & BOMANS H.E. (1997) Quelques Lucanidae de Nouvelle-Guinée dont Penichrolucanus leveri Arrow, une nouvelle espèce de Figulus MacLeay et la larve de Figulus sulcicollis Hope & Westwood (Coleoptera). Bull. Annls. Soc. R. belge Ent., 133:3-20.

ZACHER F. (1913) Die Schadlinge der Kokospalmen auf den Sudseeinseln. Arb. Kais. Biol. Aust. Land- und Forsten, 9(1):73-107.

 

Prosopocoilus (Metopodontus) bison ssp. hortensis (KRIESCHE, 1924) [Metopodontus].

Solomon Is. : Bougainville I.--6°05S-155°14E (Kieta), Shortland I.--6°55S-156°04E, Ranongga I.--8°03S-156°33E, Malaita I.--8°59S-160°59E, Gardner I.*.

 

Prosopocoilus (Metopodontus) blanchardi (PARRY, 1873) [Metopodontus, Cladognathus] [branchardi].

= Metopodontus blanchardi var. thibetanus PLANET 1899 [Thibet].

Parry, 1873:337, pl.5; Planet, 1899:385; Heyne & Taschenberg, 1908:53; Barthe, 1909:6; Miwa, 1932:127, pl.13; Didier & Séguy, 1953:41; Benesh, 1960:64; Nomura, 1965:141; Maes, 1981:3; Sakaino, 1981:23; Klausnitzer, 1982:28; Lee, Kim & Kim, 1985:417; Maes, 1990:6; Maes, 1992:70; Nomura & Lee, 1992:88; Kawano, 1993:12; Ma, 1993:278; Sakaino, Yu & Chu, 1994:123; Bartolozzi & Sforzi, 1994:114; Mizunuma & Nagai, 1994:25-26, 28, 254, pl. 73-74.

Russia : Siberia; Mongolia*;

South Korea : Cheju-do I. (Mt. Halla-san--33°23N-126°30E--Kaewol Bridge);

China : Gansu = Kansu, Beijing (Beijing), Sichuan (Kwanhsien), Hunan (Wuling Mts.), Xizang = Thibet, Yunnan, Fujian;

Nepal;

Vietnam : Tonkin (Lao Kay);

Taiwan : Taichung Distr. : Lishan

Taiwan : Hualien Distr. : Taihorin

Taiwan : Nantou Distr. : Nanshanhsi

Taiwan : Nantou Distr. : Jiyuetan

Taiwan : Nantou Distr. : Puli = Hori

Taiwan : Chiayi Distr. : Arisan

Taiwan : Kaohsiung Distr. : Liukuei

Taiwan : Tao Yuan County : Upper Pa Ling, V-1998 (3 males, 3 females in col. Ronald Cave, USA).

Taiwan : Baibara

Taiwan : Naihunpo

Taiwan : Kobayashi

Note : Mizunuma & Nagai consider this species as a ssp. of astacoides.

 

Prosopocoilus (Metopodontus) cinnamomeus (GUERIN-MENEVILLE, 1843) [Lucanus, Metopodontus].

= Lucanus pallidipennis HOPE 1841 [Java].

= Lucanus castaneus HOPE & WESTWOOD 1845 [India, error].

= Lucanus fulvipes HOPE & WESWOOD 1845 [Assam].

= Lucanus confusus HOPE & WESTWOOD 1845 [Java].

= Metopodontus javanus (FRUHSTORFER) OBERTHUR & HOULBERT 1913.

= Prosopocoilus laterinus DIDIER; KAWANO 1997.

Note : what Mizunuma & Nagai and Kawano (1997) call Prosopocoilus laterinus seems a different form but I am not shure if there are two species or two morphs in the same species.

Griffith, 1832:pl.51; Hope, 1840:77; Guerin, 1829-44 (1943):108, pl.27; Westwood, 1844:106; Hope & Westwood, 1845:12-13; Westwood, 1845:272, pl.20; Westwood, 1846:59; Burmeister, 1847:370, 372; Parry, 1870:77; Heyne & Taschenberg, 1908:53, pl.7; Boileau, 1913:224;   Oberthur & Houlbert, 1913:382, 385; Griffini, 1920:47; Didier & Séguy, 1953:40; Benesh, 1960:65; Maes, 1981:3; Sakaino, 1981:23; Weng-Yew Tung, 1983:52, pl.8; Maes, 1990:6; Maes, 1992:70; Mizunuma & Nagai, 1994:25-26, 28, 253, 254, pl. 74, 139; Kawano, 1997:454.

Java* : W. Java (Mt. Gede, Mt. Mas), Soekaboemi, Togoe, Mt. Moeria, Mt. Oeker, Mt. Tengger, Buitenzorg, Passurvan--Mt. Kawi, Tjigocka, Tjolo, Waterstand, Tjikidang, Gounod Kassi, Mt. Ardjoeno, Pengalengan, Gounod Gedeh, Dessa Tjibogo, Bukit Gompong, Znider, Malang, Mt. Smeroe.

Note : Mizunuma & Nagai consider this species as a ssp. of astacoides.

 

Prosopocoilus (Metopodontus) downesii (HOPE, 1835).

Hope & Westwood, 1845:11-12, 19; Burmeister,1847:374; Parry, 1870:81, pl.2; Griffini, 1906:138-139; Heyne & Taschenberg, 1908:53, pl.7; Boileau, 1913:223; Griffini, 1920:197; Burgeon, 1937:4; Burgeon, 1938:59; Didier & Séguy, 1952:pl.72; Basilewsky, 1956; Benesh, 1960:67, 73, 156; Maes, 1981:3; Klausnitzer, 1982:28; Bomans, 1989:443; Franciscolo, 1990:197-198; Maes, 1990:6, 9-10; Bomans, 1992:251, 252; Maes, 1992:71; Mizunuma & Nagai, 1994:240-241, pl. 55.

 

Prosopocoilus (Metopodontus) downesii ssp. downesii (HOPE, 1835).

Lucanus downesii HOPE, 1835: 99, pl.13.

Castelnau, 1840:172.

Prosopocoilus (Metopodontus) downesii ssp. downesii HOPE; Maes, 1998:280.

Bioko I.--3°31N-8°42E = Fernando Po I.* : Punta Frailes, Bahia de San Carlos, Basilé;

Principe I.--1°37N-7°23E;

Sao Tomé I.--0°13N-6°31E : Neves.

 

Prosopocoilus (Metopodontus) downesii ssp. savagei (HOPE, 1842).

Lucanus savagei HOPE, 1842.

Lucanus ungulatus HOPE, 1842:494 [Cape Palmas].

Prosopocoilus savagei ab. surtur KRIESCHE, 1920:161 [Cameroon : Buea, Barombi, Lomie, Bangwe-Land, Banyang-Land, Victoria, Moliwe bei Victoria, Nssanakang, Bibundi, Akonolingo-Abong Mbang, Lolodorf, Ossidinge] (could be a ssp. from Cameroon-Zaire).

Prosopocoilus savagei ab. metaxylimnus KRIESCHE, 1920:161 [Victoria Lake : Ukerewe I.].

Prosopocoilus (Metopodontus) downesii ssp. savagei (HOPE); Maes, 1998:280.

Senegal

Guinea : Mt. Nimba : Kéoulenta, 7°43 N - 8°20 W

Sierra Leone : N. Prov. : Bumbuna

Liberia : Cape Palmas*--4°23N-7°36W, Junk River;

Ivory Coast : Dimbokro--6°41N-4°34W, Bingerville, Divo--5°56N-5°29W, Eloka, Tiassale--5°59N-4°52W, Abidjan--5°19N-4°02W;

Ghana : Accra--5°35N-0°16W;

Togo : Bismarckburg--8°11N-0°39E;

Benin (Dahomey);

Nigeria = Ashanti = Calabar = Old Calabar;

Cameroon : Isabo, Fumsam, Lolodorf--3°12N-10°45E, Samba, Edea--3°47N-10°07E, Yaoundé--3°51N-11°30E, Victoria (Moliwe), Buea--4°11N-9°20E, Barombi, Lomie, Bangwe, Banyang-Land, Nssanakang, Bibundi, Akonolingo--Mbang, Ossidinge;

Central African Republic;

Equatorial Guinea

Gabon :

Petit-Okano, 0°02 S - 11°53 E, IX-85, leg. J.M. Weill & A. Pauly (in coll. Musée royal de l'Afrique centrale, Bruxelles, Belgique).

Petit-Okano, 0°02 S - 11°53 E, III-86, leg. A. Pauly (in coll. Musée royal de l'Afrique centrale, Tervuren, Belgique).

Gabon : Libreville--0°28N-9°25E (in coll. Maes, Nicaragua).

Gabon, ex coll. J. Carradot (in coll. Institut royal des Sciences naturelles de Belgique).

Gabon, ex coll. Steinmetz (in coll. Institut royal des Sciences naturelles de Belgique).

Gabon français, ex coll. Le Moult (in coll. Institut royal des Sciences naturelles de Belgique).

Gabon (in coll. Maes, Nicaragua).

Congo : Mougoumba, Kintele, Sibiti, Mbila, Dimonika--4°14S-12°25E, Odouka;

Zaire : Haut-Zaire [Albert Lake--1°38N-30°52E = Maputo Lake (Mboga, West-Lendu, Wabudso-Wald--Wakangu), Albert National Park (Kibga--S. Mt. Visoke), Beni--0°29N-29°46E, Bunia--1°39N-30°26E, Rwankwi, Moku--2°31N-25°58E], Kivu [Mohangi, Mulungu, Kisantu], Kasai [Dekese--3°33S-21°13E, Lusambo--5°01S-23°25E, Sankuru River (Kondue)], Bas-Zaire [Maduda--4°55S-13°06E], ? [Voka];

Rwanda : Bulera Lake--1°26S-29°47E (Kagogo);

Uganda : Budongo Forest--1°46N-31°33E;

Tanzania : Mwanza [Victoria Lake (Ukerewe I.--2°00S-32°58E)].

Xylophagous : Moraceae : Castilla.

Bibliography

MAES J.M. (1998) Lucanidae (Coleoptera) du Gabon. Bull. Annls. Soc. r. Belge Ent., 134(1998):279-285.

 

Prosopocoilus (Metopodontus) dubernardi (PLANET, 1899) [Metopodontus].

Planet, 1899:35; Didier & Séguy, 1953:40; Benesh, 1960:67; Maes, 1981:3; Klausnitzer, 1982:43; Maes, 1990:6; Maes, 1992:71; Mizunuma & Nagai, 1994:25-26, 28, 254, pl. 73.

China: Sichuan = Sseutchouan, Xizang = Thibet (Tsa Kou*, Manang-Kouang), Yunnan (Yunnan-Sen, Kouang-Si-Hien, Djo Kou La - 31°25 N - 102°58 E, Pe-Yen-Tsing).

Note : Mizunuma & Nagai consider this species as a ssp. of astacoides.

 

Prosopocoilus (Metopodontus) elaphus (MOLLENKAMP, 1902) [Metopodontus].

= Prosopocoilus diehli WEINREICH 1971 [Metopodontus] [Sumatra : Tebingtinggi].

Mollenkamp, 1902:283; Mollenkamp, 1903:342; Benesh, 1960:67; Weinreich, 1971:230, pl.7; Maes, 1981:3, 4; Weng-Yew Tung, 1983:52, pl.8; Maes, 1990:6; Maes, 1992:71; Mizunuma & Nagai, 1994:25-26, 28, 254, pl. 74; Kawano, 1997:454.

Malaya : Perak (Taiping--4°51N-100°41E, Kampong Sahom), Pahang (Cameron Highlands);

Sumatra : N. Sumatra Prov. (Doloc Baros Estate--Medan, Toba*, Tebingtinggi, Berastagi), Nias I., W. Sumatra Prov. (Solok), S. Sumatra Prov. (Mt. Dempo).

Note : Mizunuma & Nagai consider this species as a ssp. of astacoides.

 

Prosopocoilus (Metopodontus) formosus (LACROIX) [Metopodontus].

Lacroix; Bomans, 1991:146.

Note : Pierre Benoit (in litteris) consider this as a synonym of Prospocoilus astacoides.

Thailand : Chiang Mai (Mt. Doi Pui), Tak (road Mae-Sot--Umphang).

 

Prosopocoilus (Metopodontus) kannegieteri (VAN DE POLL, 1895) [Metopodontus].

Metopodontus kannegieteri VAN DE POLL, 1895:63 (Sumatra: Ranau).

Metopodontus volkmanni NONFRIED, 1895:279 [Sumatra : Siboga].

Ritsema, 1896:130; Benesh, 1960:69

Prosopocoilus kannegieteri subsp. obscurus WEINREICH, 1971:236, pl.9 [Sumatra : Langkat--Balei Gadjah, Medan--Dolok Baros Estate].

Klausnitzer, 1982:43; Maes, 1990:6; Bomans, 1991:146; Maes, 1992:71

Prosopocoilus kannegieteri (VAN DE POLL); Mizunuma & Nagai, 1994: 252, pl. 71, fig.325(1-11).

Metopodontus kannegieteri VAN DE POLL; Bomans & Benoit, 2005:293.

Ø      Thailand : ? : Suratani

Ø      Malaya : Kedah, IV-1988 (1 female in coll. Mizunuma, Japan).

Ø      Malaya : Perak : Taiping, 4°51N-100°41E

Ø      Malaya : Pahang : Cameron Highlands, IV-1992 (1 male in coll. Mizunuma, Japan).

Ø      Borneo : Sabah : Crocker Range, IV-1983 (2 males in coll. Mizunuma, Japan).

Ø      Borneo : Sabah : Crocker Range, III-1990 (1 male in coll. Mizunuma, Japan).

Ø      Borneo : Sabah : Crocker Range, 27-V-1992 (1 female in coll. Mizunuma, Japan).

Ø      Sumatra : N. Sumatra Prov. : Doloc Baros Estate : Medan

Ø      Sumatra : S. Sumatra Prov. : Palembang Residency : Mts. near lake Ranau, 3000 Ft., IV-1890, col. J.Z. Kannegieter (male TYPUS in Rijksmuseum van Naturlijke Historie, Leiden, Holland).

Ø      Sumatra : Lampung Prov., III-1987 (2 males, 1 female in coll. Mizunuma, Japan).

Ø      Sumatra : ? : Balei Gadjah : Langkat

Ø      Sumatra : ? : Siboga

Ø      Sumatra : ? : Si Gawak

Ø      Philippines : Palawan I.--9°37N--118°32E, VII-1992 (1 male in coll. Mizunuma, Japan).

Bibliography.

BOMANS H.E. & BENOIT P. (2005) Inventaire provisoire commenté des Coléoptères Lucanides de l’Archipel des Philippines (98eme Contribution a l’étude des Coléoptères Lucanides). Lambillionea, CV(2):283-301.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Prosopocoilus (Metopodontus) mirabilis (BOILEAU, 1904) [Metopodontus].

Boileau, 1904:278; Kriesche, 1920:162; Benesh, 1960:70; Maes, 1981:3; Maes, 1990:6; Maes, 1992:71.

Tanzania : Tanga [Usambara*], Pwani [Bagamoyo--6°30S-38°55E], Pwani [Dar Es Salaam].

 

Prosopocoilus (Metopodontus) torresensis (DEYROLLE, 1870) [Metopodontus, Prosopocoelus].

= Cladognathus limbatus WATERHOUSE 1887 [Metopodontus, Prosopocoilus, Prosopocoelus] [Torres Straits].

= Metopodontus torresensis var. flavoguttatus MOLLENKAMP 1906 [Prosopocoilus] [Australia : Queensland : Endeavour River].

Deyrolle en Parry, 1870:80, pl.1; Gestro, 1881:306; Waterhouse, 1887:381; Van Roon, 1905:117; Mollenkamp, 1906:31; Van Roon, 1910:23; Boileau, 1913:225; Arrow, 1915:502; Kriesche, 1924:439; Nagel, 1930:91; Didier & Séguy, 1953:106; Benesh, 1960:64, 75; Maes, 1981:4; Klausnitzer, 1982:43; Lacroix, 1988:12; Maes, 1990:6; Maes, 1992:71; Moore & Cassis, 1992:15; Mizunuma & Nagai, 1994:245, pl. 61; Wood, Hasenpusch & Storey, 1996:47.

Australia : Northern Territory (Fergusson River), Queensland (Mt. Cornwallis--9°25S-142°31E, Murray I.--9°56S-144°04E, Somerset--10°45S-142°35E, Endeavour River--15°22S-145°02E, Daintree--16°15S-145°19E, Kuranda--16°49S-145°38E, Cairns--16°55S-145°46E, Mt. French--28°00S-152°37E, Cape York--33°15S-150°08E, Gardner I., Toeal), Torres Straits Is.* (Thursday I.--10°35S-142°13E, Tawan I.--22°10S-152°47E); ? Victoria.

Saproxylophagous.

 

Prosopocoilus (Metopodontus) umhangi (FAIRMAIRE, 1891) [Cladognathus, Metopodontus].

= Prosopocoilus hacquarti OBERTHUR 1880 [Bagamoyo].

= Metopodontus wentzel-heckmannae KOLBE 1901 [Annam, error].

= Metopodontus elisabethae WATERHOUSE 1905 [German East Africa : Usagara].

Oberthur, 1880:cxix; Fairmaire, 1891:cxcii; Planet, 1894:119; Planet, 1899:388, pl.1; Kolbe, 1901:71; Waterhouse, 1905:375; Zang, 1906:94; Felsche, 1907:146; Kriesche, 1920:163; Didier & Séguy, 1952:pl.69; Didier & Séguy, 1953:42; Benesh, 1960:75; Maes, 1981:3; Klausnitzer, 1982:28; Bomans, 1989:443; Maes, 1990:6; Maes, 1992:71; Mizunuma & Nagai, 1994:240, pl. 54.

Tanzania = German East Africa* : Tanga [Usambara (Bulwa, Amani)], Dodoma [Usagara region--6°45S-36°40E], Morogoro [Mhonda--6°13S-37°37E, Usegua region--6°20S-37°45E, Nguru Mts.], Pwani [Bagamoyo--6°30S-38°55E], Mtwara [Lindi*--10°02S-39°39E];

Zanzibar I.--6°09S-39°14E;

Malawi = Nyassa.


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