Hexarthrius HOPE, 1843

(G. = Hexarthrius parryi)


Hexarthrius aduncus JORDAN, 1894.

Jordan, 1894:484, pl.13; Arrow, 1950:73, pl.6; Didier & Séguy, 1952:pl.29; Benesh, 1953:25; Benesh, 1960:128; Maes, 1992:21; Kawano, 1993:12; Mizunuma & Nagai, 1994:259, pl. 85.

India : Assam*, Meghalaya (Khasi Hills, Shillong), Manipur.

 

Hexarthrius bowringii PARRY, 1862 [bowringi].

Parry, 1862:108; Parry, 1864:12, pl.9; Arrow, 1950:72, pl.6; Didier & Séguy, 1952:pl.28; Benesh, 1960:128; Maes, 1992:21; Mizunuma & Nagai, 1994:258, pl. 84, 139.

India* : West Bengal (Darjeeling), Assam;

Myanmar = Burma.

 

Hexarthrius buquettii (HOPE, 1843) [Lucanus] [buqueti] [Philippines : Manilla, error].

= Lucanus falciger HOPE & WESTWOOD 1845 [Java].

= Lucanus longipennis HOPE & WESTWOOD 1845 [Java].

= Lucanus rhinoceros BURMEISTER (nec OLIVIER) 1847 [Java].

Hope, 1843:95; Hope & Westwood, 1845:10, 11; Hope, 1846:182, pl.13; Burmeister, 1847:366; Reiche, 1853:75; Griffini, 1906:899; Heyne & Taschenberg, 1908:50, pl.6; Boileau, 1913:221; Oberthur & Houlbert, 1913:273; Griffini, 1918:114; Didier & Séguy, 1952:pl.31; Benesh, 1960:128; Klausnitzer, 1982:42; Maes, 1992:21; Kawano, 1993:13; Mizunuma & Nagai, 1994:258, pl. 84; Kawano, 1997:454.

Java : W. Java : Mt. Gede

Java : W. Java : Mt. Mas

Java : W. Java : Sukabumi : Mt. Halimun, 800-1200 m, V-2000, leg. A. Hasan (1 male in col. Ronald Cave, USA).

Java : E. Java : Mt. Arjuna

Java : Tjikidang

Java : Tjolo : Mt. Moeria

Java : Tjigoska

Java : Soekaboemi

Java : Toegoe

Java : Waterstrand

Java : Mt. Kawie

Java : Malang

Java : Mt. Ardjoeno

Java : Mt. Salak.

 

Hexarthrius davisoni WATERHOUSE, 1888 [Borneo : Kinabalu, error].

= Hexarthrius cotesi NONFRIED 1892 [India].

= Hexarthrius castetsi BOILEAU 1897 [Inde Or.].

Waterhouse, 1888:260; Nonfried, 1892:365; Boileau, 1897:581; Boileau, 1898:411; Planet, 1899:174; Gravely, 1915:414; Arrow, 1950:74, pl.6; Didier & Séguy, 1952:pl.30; Benesh, 1953:26; Benesh, 1960:128; Klausnitzer, 1982:18; Arif, Shah & Joshi, 1990:36; Arif, Shah & Joshi, 1991:73-74; Maes, 1992:21; Mizunuma & Nagai, 1994:259, pl. 85.

India : Tamil Nadu (Madras, Madurai, Kodaikanal, Madurai--Shambaganur, Anamalai Hills, Palni Hills, Travancore, Erichinopolis, Cuddapah, Pulney Hills).

Note : Arif, Shah & Joshi (1990-1991) cites this species from Himalaya : Uttar Pradesh : Joshimath, this locality needs verification.

Saproxylophagous :

Fagaceae : Quercus.

Juglandaceae : Juglans.

? : pangar.

Adults on Fagaceae :

Quercus.

Juglandaceae : Juglans.

? : pangar.

Fungivorous : Pleurotus.

 

Hexarthrius forsteri (HOPE, 1840) [Lucanus].

= Lucanus serricollis HOPE 1840 [Assam].

Hope, 1840:77-78; Hope, 1841:591, pl.40; Hope, 1841:299; Hope & Westwood, 1845:11; Burmeister, 1847:367; Parry, 1863:446; Heyne & Taschenberg, 1908:51; Boileau, 1913:222; Gravely, 1915:414; Arrow, 1950:69, pl.5-6; Didier & Séguy, 1952:pl.36; Benesh, 1960:129; Klausnitzer, 1982:18; Maes, 1992:21; Mizunuma & Nagai, 1994:258, pl.             84.

India : Assam*, Meghalaya (Khasi Hills, Shillong, Jaintai Hills);

Bhutan;

Cambodia (Kep sur Mer).

 

Hexarthrius howdeni DE LISLE, 1972.

Hexarthrius howdeni DE LISLE, 1972:151 (Calapan).

Maes, 1992:21

Hexarthrius howdeni DE LISLE; Mizunuma & Nagai, 1994:258, pl. 82, 139, fig.342(1-3).

Hexarthrius howdeni DE LISLE; Bomans & Benoit, 2005:292.

Ø      Philippines : Luzon I. : Mountain Province, XII-1987 (1 male in coll. Mizunumai, Japan).

Ø      Philippines : Mindoro I. : San Luis, Calapan, 13°24N-121°11E, 13-V-1954, leg. David Towns (male TYPUS in coll. Howden, Canada).

Ø      Philippines : Mindoro I. : Mt. Halcon, II-1991 (1 male in coll. Mizunumai, Japan).

Ø      Philippines : Mindoro I., V-1992 (1 female in coll. Mizunumai, Japan).

Bibliography.

BOMANS H.E. & BENOIT P. (2005) Inventaire provisoire commenté des Coléoptères Lucanides de l’Archipel des Philippines (98eme Contribution a l’étude des Coléoptères Lucanides). Lambillionea, CV(2):283-301.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Hexarthrius mandibularis ssp. mandibularis DEYROLLE, 1881.

Hexarthrius mandibularis DEYROLLE, 1881:237, pl. 5 [Borneo] [mundibularis].

Nagel, 1926:293; Nagel, 1927:145; De Jong, 1937:553; Didier & Séguy, 1952:pl.37; Benesh, 1960:129; Weng-Yew Tung, 1983:49, pl.7; Kawano, 1988:26; Maes, 1992:21; Kawano, 1993:12.

Hexarthrius mandibularis ssp. mandibularis DEYROLLE; Mizunuma & Nagai, 1994:26, fig. 79(a-c), p. 258, pl. 83-84, 139.

Kawano, 1997:454.

Borneo (TYPUS).

Borneo : Sarawak : Mt. Penrisson.

Borneo : Sarawak : Kuching : Mt. Serapi

Borneo : Sabah : Crocker Range.

 

Hexarthrius mandibularis ssp. sumatranus MIZUNUMA, 1994.

Hexarthrius mandibularis ssp. sumatranus MIZUNUMA in MIZUNUMA & NAGAI, 1994:26, 28, fig. 80(a-c), pl. 84, fig. 344(8-13) [Lampung Prov.].

Sumatra : W. Sumatra Prov. : Padangpanjang, IV-1984 (1 male PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : S. Sumatra Prov. : Palembang

Sumatra : Lampung Prov., III-1986 (1 male PARATYPUS in col. H. Fujita, Japan).

Sumatra : Lampung Prov., IX-1986 (male HOLOTYPUS in Kanagawa Prefectural Museum, Japan) (12 males PARATYPUS, 1 female PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : Lampung Prov., IV-1992 (25 males PARATYPUS in col. H. Fujita, Japan).

Sumatra : Lampung Prov., V-1993 (2 males PARATYPUS in col. H. Fujita, Japan).

Sumatra : Lampung Prov., V/VI-1993 (30 males PARATYPUS, 1 female PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : Deli.

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Hexarthrius melchioritis SEGUY, 1954.

Séguy, 1954:269; De Lisle, 1975:267; Benesh, 1960:157; Maes, 1992:21.

Myanmar = Burma : Kachin State (Kachin Kauri*).

 

Hexarthrius mniszechi (THOMSON, 1857) [Lucanus] [mniszechii, mniszecki].

Lacordaire, 1856:pl.24; Thomson, 1857:396; Gravely, 1915:414; Arrow, 1950:71, pl.6; Didier & Séguy, 1952:pl.35; Benesh, 1960:129; Maes, 1992:22; Bartolozzi & Sforzi, 1994:114; Mizunuma & Nagai, 1994:258, pl. 84.

India : Assam (Sibsagar), Meghalaya (Khasi Hills);

Nepal;

Bangladesh : Sylhet*;

Myanmar = Burma : Kachin State (Kauri--Kachin Hills).

 

Hexarthrius nigritus LACROIX, 1990.

Lacroix, 1990:12-13, pl.1; Bomans, 1991:146; Kawano, 1992:11; Mizunuma & Nagai, 1994:257, pl. 82; Kawano, 1997:454.

Thailand : Chiang Mai (Chiang Mai--18°50N-98°57E), Saraburi (Khao Yai--14°30N, Saraburi), Nakhon Ratchasima (Nakhon Ratchasima--15°01N-102°04E), ? (Tak Mas Song*).

 

Hexarthrius parryi HOPE, 1842.

Hope, 1842:127; Hope, 1842:247; Hope, 1843:104, pl.10; Burmeister, 1847:367; Parry, 1864:11, pl.4; Jordan, 1894:484; Mollenkamp, 1897:146; Boileau, 1904:277; Mollenkamp, 1904:402; Mollenkamp, 1905:7; Heyne & Taschenberg, 1908:50, pl.6; Gravely, 1915:414; Arrow, 1950:68, pl.5-6; Didier & Séguy, 1952:pl.32; Benesh, 1960:129; Bomans, 1970:225; Weinreich, 1971; Weinreich, 1972; Klausnitzer, 1982:14, 42; Weng-Yew Tung, 1983:48, pl.4; Kawano, 1988:22; Lacroix, 1990:12-13,?91; Maes, 1990:2; Téllez, 1990:39; Bomans, 1991:146; Maes, 1992:21-22; Kawano, 1992:11; Bomans, 1994:493; Kawano, 1994:9, 11; Mizunuma & Nagai, 1994:257, 258, pl. 81-82, 139; Kawano, 1997:454.

Hexarthrius parryi ssp. parryi HOPE, 1842.

India : Sikkim, Assam (Cachar, Sibsagar), Meghalaya (Shillong, Khasi Hills), Nagaland (Kohima);

Bangladesh : Sylhet*;

Myanmar = Burma;

Thailand;

Laos : Sayabouri;

Cambodia : Kep sur Mer;

Vietnam : Tonkin.

Hexarthrius parryi ssp. deyrollei PARRY, 1864 [deyrollii].

Myanmar = Burma : Shan State (Muang Yoo), Tenasserim State;

Thailand = Siam* : Chiang Rai (Chiang Rai, Wiangpapao, Mae Jan, Maekajan), Chiang Mai (Fang);

Malaya : Perak (Taiping--4°51N-100°41E, Kampong Sahom), Pahang (Cameron Highlands).

Hexarthrius parryi ssp. paradoxus MOLLENKAMP, 1897.

Sumatra* : N. Sumatra Prov. (Medan--Doloc Baros, Toba Lake, Berastagi), W. Sumatra Prov. (Padang), Lampung Prov., ? (Langkat--Balei Gadjah, Sipirok);

Java.

Hexarthrius parryi ssp. elongatus JORDAN, 1894.

= Hexarthrius rollei MOLLENKAMP 1904 [Kinabalu].

= Hexartrius deyrollei var. rufipes BOILEAU 1904 [Kinabalu].

Borneo : Sabah (Mt. Kinabalu*, Crocker Range), W. Kalimantan (Mt. Bawang near Singkawang).

 

Hexarthrius rhinoceros ssp. rhinoceros (OLIVIER, 1789).

Lucanus rhinoceros OLIVIER, 1789:21-22, pl. 5 [Amérique méridionale].

Lucanus vitulus DEJEAN, 1837:193 [nomen nudum] [Java].

Olivier (ed. Illiger), 1800:73; Olivier (ed. Sturm), 1802:6, pl.4; Thunberg, 1806:201; Hope, 1843:95; Hope, 1845:183; Burmeister, 1847:366; Reiche, 1853:75; Deyrolle, 1864:312, pl.4; Vollenhoven, 1865:148; Parry, 1870:56; Planet, 1899:47; Heyne & Taschenberg, 1908:50; Oberthur & Houlbert, 1913:218; Didier & Séguy, 1952:pl.33-34; Benesh, 1960:129-130; Maes, 1992:22; Kawano, 1993:12.

Hexarthrius rhinoceros ssp. rhinoceros (OLIVIER); Mizunuma & Nagai, 1994:26, fig. 76(a-b), p. 258, pl. 83, 139.

Java : W. Java : Mt. Halimun.

Java : W. Java : Mt. Gede.

Java : E. Java : Mt. Kawi.

Java : Wangoen-Tjisampara.

 

Hexarthrius rhinoceros ssp. chaudoiri DEYROLLE, 1864.

Hexarthrius chaudoiri DEYROLLE, 1864 [Sumatra].

Hexarthrius rhinoceros ssp. chaudoiri DEYROLLE; Mizunuma & Nagai, 1994:26, 28, fig. 77(a-b), p. 258, pl. 83, 139.

Sumatra (TYPUS).

Sumatra : W. Sumatra Prov. : Padang.

Sumatra : W. Sumatra Prov. : Bukittingi.

 

Hexarthrius rhinoceros ssp. sadaoi MIZUNUMA, 1994.

Hexarthrius rhinoceros ssp. sadaoi MIZUNUMA in MIZUNUMA & NAGAI, 1994:26, 28, fig. 78(a-b), pl. 139, fig. 343(8-9) [Pagai I.].

Sumatra : W. Sumatra Prov. : Mentawei Is. : Siberut I., IV-1993 (2 males PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : W. Sumatra Prov. : Mentawei Is. : Sipora I.

Sumatra : W. Sumatra Prov. : Mentawei Is. : Pagai I., II-1993 (male HOLOTYPUS in Kanagawa Prefectural Museum, Japan) (3 males PARATYPUS in col. T. Mizunuma, Japan).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Hexarthrius vitalisi DIDIER, 1925.

Didier, 1925:202; Didier & Séguy, 1953:89; Benesh, 1960:130; Bomans, 1991:146; Maes, 1992:22; Kawano, 1993:12; Kawano, 1994:9; Mizunuma & Nagai, 1994:259, pl. 85; Kawano, 1997:454.

Thailand : Chiang Rai (Wiangpapao), Chiang Mai (Chiang Mai--18°50N-98°57E), Lampang (Muang Pan);

Vietnam : Tonkin (Tam Dao near Vinh Yen, Paklay*, Chapa*).


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