Hexarthrius HOPE, 1843
(G. = Hexarthrius parryi)
Hexarthrius aduncus JORDAN, 1894.
Jordan, 1894:484, pl.13; Arrow, 1950:73,
pl.6; Didier & Séguy, 1952:pl.29; Benesh, 1953:25; Benesh, 1960:128; Maes,
1992:21; Kawano, 1993:12; Mizunuma & Nagai, 1994:259, pl. 85.
India :
Assam*, Meghalaya (Khasi Hills, Shillong), Manipur.
Hexarthrius
bowringii PARRY, 1862 [bowringi].
Parry,
1862:108; Parry, 1864:12, pl.9; Arrow, 1950:72, pl.6; Didier & Séguy,
1952:pl.28; Benesh, 1960:128; Maes, 1992:21; Mizunuma & Nagai, 1994:258,
pl. 84, 139.
India* :
West Bengal (Darjeeling), Assam;
Myanmar =
Burma.
Hexarthrius
buquettii (HOPE, 1843) [Lucanus] [buqueti]
[Philippines : Manilla, error].
= Lucanus
falciger HOPE & WESTWOOD 1845 [Java].
= Lucanus
longipennis HOPE & WESTWOOD 1845 [Java].
= Lucanus rhinoceros BURMEISTER (nec
OLIVIER) 1847 [Java].
Hope, 1843:95; Hope & Westwood, 1845:10,
11; Hope, 1846:182, pl.13; Burmeister, 1847:366; Reiche, 1853:75; Griffini,
1906:899; Heyne & Taschenberg, 1908:50, pl.6; Boileau, 1913:221; Oberthur
& Houlbert, 1913:273; Griffini, 1918:114; Didier & Séguy, 1952:pl.31;
Benesh, 1960:128; Klausnitzer, 1982:42; Maes, 1992:21; Kawano, 1993:13;
Mizunuma & Nagai, 1994:258, pl. 84; Kawano, 1997:454.
Java : W. Java : Mt. Gede
Java : W. Java : Mt. Mas
Java : W. Java : Sukabumi : Mt. Halimun,
800-1200 m, V-2000, leg. A. Hasan (1 male in col. Ronald Cave, USA).
Java : E. Java : Mt. Arjuna
Java : Tjikidang
Java : Tjolo : Mt. Moeria
Java : Tjigoska
Java : Soekaboemi
Java :
Toegoe
Java :
Waterstrand
Java : Mt. Kawie
Java : Malang
Java : Mt. Ardjoeno
Java : Mt. Salak.
Hexarthrius
davisoni WATERHOUSE, 1888 [Borneo : Kinabalu, error].
= Hexarthrius
cotesi NONFRIED 1892 [India].
= Hexarthrius
castetsi BOILEAU 1897 [Inde Or.].
Waterhouse,
1888:260; Nonfried, 1892:365; Boileau, 1897:581; Boileau, 1898:411; Planet,
1899:174; Gravely, 1915:414; Arrow, 1950:74, pl.6; Didier & Séguy,
1952:pl.30; Benesh, 1953:26; Benesh, 1960:128; Klausnitzer, 1982:18; Arif, Shah
& Joshi, 1990:36; Arif, Shah & Joshi, 1991:73-74; Maes, 1992:21;
Mizunuma & Nagai, 1994:259, pl. 85.
India :
Tamil Nadu (Madras, Madurai, Kodaikanal, Madurai--Shambaganur, Anamalai Hills,
Palni Hills, Travancore, Erichinopolis, Cuddapah, Pulney Hills).
Note :
Arif, Shah & Joshi (1990-1991) cites this species from Himalaya : Uttar
Pradesh : Joshimath, this locality needs verification.
Saproxylophagous :
Fagaceae : Quercus.
Juglandaceae
: Juglans.
? :
pangar.
Adults on
Fagaceae :
Quercus.
Juglandaceae : Juglans.
? : pangar.
Fungivorous
: Pleurotus.
Hexarthrius
forsteri (HOPE, 1840) [Lucanus].
= Lucanus
serricollis HOPE 1840 [Assam].
Hope,
1840:77-78; Hope, 1841:591, pl.40; Hope, 1841:299; Hope & Westwood,
1845:11; Burmeister, 1847:367; Parry, 1863:446; Heyne & Taschenberg,
1908:51; Boileau, 1913:222; Gravely, 1915:414; Arrow, 1950:69, pl.5-6; Didier
& Séguy, 1952:pl.36; Benesh, 1960:129; Klausnitzer, 1982:18; Maes, 1992:21;
Mizunuma & Nagai, 1994:258, pl. 84.
India :
Assam*, Meghalaya (Khasi Hills, Shillong, Jaintai Hills);
Bhutan;
Cambodia (Kep sur Mer).
Hexarthrius howdeni DE LISLE, 1972.
Hexarthrius howdeni DE LISLE, 1972:151 (Calapan).
Maes, 1992:21
Hexarthrius
howdeni DE LISLE; Mizunuma & Nagai,
1994:258, pl. 82, 139, fig.342(1-3).
Hexarthrius
howdeni DE LISLE; Bomans & Benoit, 2005:292.
Ø Philippines : Luzon
I. : Mountain Province, XII-1987 (1 male in coll. Mizunumai, Japan).
Ø Philippines : Mindoro I. : San Luis, Calapan, 13°24N-121°11E,
13-V-1954, leg. David Towns (male TYPUS in coll. Howden, Canada).
Ø Philippines : Mindoro
I. : Mt. Halcon, II-1991 (1 male in coll. Mizunumai, Japan).
Ø Philippines : Mindoro
I., V-1992 (1 female in coll. Mizunumai, Japan).
Bibliography.
BOMANS
H.E. & BENOIT P. (2005) Inventaire provisoire commenté des Coléoptères
Lucanides de l’Archipel des Philippines (98eme Contribution a l’étude des
Coléoptères Lucanides). Lambillionea, CV(2):283-301.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Hexarthrius
mandibularis ssp. mandibularis DEYROLLE, 1881.
Hexarthrius
mandibularis DEYROLLE, 1881:237, pl. 5 [Borneo] [mundibularis].
Nagel,
1926:293; Nagel, 1927:145; De Jong, 1937:553; Didier & Séguy, 1952:pl.37; Benesh,
1960:129; Weng-Yew Tung, 1983:49, pl.7; Kawano, 1988:26; Maes, 1992:21; Kawano,
1993:12.
Hexarthrius
mandibularis ssp. mandibularis DEYROLLE; Mizunuma & Nagai, 1994:26, fig.
79(a-c), p. 258, pl. 83-84, 139.
Kawano,
1997:454.
Borneo
(TYPUS).
Borneo :
Sarawak : Mt. Penrisson.
Borneo :
Sarawak : Kuching : Mt. Serapi
Borneo :
Sabah : Crocker Range.
Hexarthrius
mandibularis ssp. sumatranus MIZUNUMA, 1994.
Hexarthrius
mandibularis ssp. sumatranus MIZUNUMA in MIZUNUMA & NAGAI, 1994:26, 28,
fig. 80(a-c), pl. 84, fig. 344(8-13) [Lampung Prov.].
Sumatra : W. Sumatra Prov. : Padangpanjang,
IV-1984 (1 male PARATYPUS in col. T. Mizunuma, Japan).
Sumatra : S. Sumatra Prov. : Palembang
Sumatra :
Lampung Prov., III-1986 (1 male PARATYPUS in col. H. Fujita, Japan).
Sumatra :
Lampung Prov., IX-1986 (male HOLOTYPUS in Kanagawa Prefectural Museum, Japan)
(12 males PARATYPUS, 1 female PARATYPUS in col. T. Mizunuma, Japan).
Sumatra : Lampung Prov., IV-1992 (25 males
PARATYPUS in col. H. Fujita, Japan).
Sumatra : Lampung Prov., V-1993 (2 males
PARATYPUS in col. H. Fujita, Japan).
Sumatra : Lampung Prov., V/VI-1993 (30 males
PARATYPUS, 1 female PARATYPUS in col. T. Mizunuma, Japan).
Sumatra : Deli.
Bibliography.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Hexarthrius
melchioritis SEGUY, 1954.
Séguy, 1954:269; De Lisle, 1975:267; Benesh,
1960:157; Maes, 1992:21.
Myanmar =
Burma : Kachin State (Kachin Kauri*).
Hexarthrius
mniszechi (THOMSON, 1857) [Lucanus] [mniszechii,
mniszecki].
Lacordaire,
1856:pl.24; Thomson, 1857:396; Gravely, 1915:414; Arrow, 1950:71, pl.6; Didier
& Séguy, 1952:pl.35; Benesh, 1960:129; Maes, 1992:22; Bartolozzi &
Sforzi, 1994:114; Mizunuma & Nagai, 1994:258, pl. 84.
India :
Assam (Sibsagar), Meghalaya (Khasi Hills);
Nepal;
Bangladesh
: Sylhet*;
Myanmar =
Burma : Kachin State (Kauri--Kachin Hills).
Hexarthrius
nigritus LACROIX, 1990.
Lacroix, 1990:12-13,
pl.1; Bomans, 1991:146; Kawano, 1992:11; Mizunuma & Nagai, 1994:257, pl.
82; Kawano, 1997:454.
Thailand
: Chiang Mai (Chiang Mai--18°50N-98°57E), Saraburi (Khao Yai--14°30N,
Saraburi), Nakhon Ratchasima (Nakhon Ratchasima--15°01N-102°04E), ? (Tak Mas
Song*).
Hexarthrius
parryi HOPE, 1842.
Hope,
1842:127; Hope, 1842:247; Hope, 1843:104, pl.10; Burmeister, 1847:367; Parry,
1864:11, pl.4; Jordan, 1894:484; Mollenkamp, 1897:146; Boileau, 1904:277;
Mollenkamp, 1904:402; Mollenkamp, 1905:7; Heyne & Taschenberg, 1908:50,
pl.6; Gravely, 1915:414; Arrow, 1950:68, pl.5-6; Didier & Séguy,
1952:pl.32; Benesh, 1960:129; Bomans, 1970:225; Weinreich, 1971; Weinreich,
1972; Klausnitzer, 1982:14, 42; Weng-Yew Tung, 1983:48, pl.4; Kawano, 1988:22;
Lacroix, 1990:12-13,?91; Maes, 1990:2; Téllez, 1990:39; Bomans, 1991:146; Maes,
1992:21-22; Kawano, 1992:11; Bomans, 1994:493; Kawano, 1994:9, 11; Mizunuma
& Nagai, 1994:257, 258, pl. 81-82, 139; Kawano, 1997:454.
Hexarthrius
parryi ssp. parryi HOPE, 1842.
India :
Sikkim, Assam (Cachar, Sibsagar), Meghalaya (Shillong, Khasi Hills), Nagaland
(Kohima);
Bangladesh
: Sylhet*;
Myanmar =
Burma;
Thailand;
Laos :
Sayabouri;
Cambodia : Kep sur Mer;
Vietnam :
Tonkin.
Hexarthrius
parryi ssp. deyrollei PARRY, 1864 [deyrollii].
Myanmar =
Burma : Shan State (Muang Yoo), Tenasserim State;
Thailand
= Siam* : Chiang Rai (Chiang Rai, Wiangpapao, Mae Jan, Maekajan), Chiang Mai
(Fang);
Malaya :
Perak (Taiping--4°51N-100°41E, Kampong Sahom), Pahang (Cameron Highlands).
Hexarthrius
parryi ssp. paradoxus MOLLENKAMP, 1897.
Sumatra* : N. Sumatra Prov. (Medan--Doloc
Baros, Toba Lake, Berastagi), W. Sumatra Prov. (Padang), Lampung Prov., ?
(Langkat--Balei Gadjah, Sipirok);
Java.
Hexarthrius
parryi ssp. elongatus JORDAN, 1894.
= Hexarthrius
rollei MOLLENKAMP 1904 [Kinabalu].
= Hexartrius
deyrollei var. rufipes BOILEAU 1904 [Kinabalu].
Borneo :
Sabah (Mt. Kinabalu*, Crocker Range), W. Kalimantan (Mt. Bawang near
Singkawang).
Hexarthrius rhinoceros ssp. rhinoceros
(OLIVIER, 1789).
Lucanus rhinoceros OLIVIER, 1789:21-22, pl. 5 [Amérique méridionale].
Lucanus vitulus DEJEAN, 1837:193 [nomen nudum] [Java].
Olivier (ed. Illiger), 1800:73; Olivier (ed.
Sturm),
1802:6, pl.4; Thunberg, 1806:201; Hope, 1843:95; Hope, 1845:183; Burmeister,
1847:366; Reiche, 1853:75; Deyrolle, 1864:312, pl.4; Vollenhoven, 1865:148;
Parry, 1870:56; Planet, 1899:47; Heyne & Taschenberg, 1908:50; Oberthur
& Houlbert, 1913:218; Didier & Séguy, 1952:pl.33-34; Benesh,
1960:129-130; Maes, 1992:22; Kawano, 1993:12.
Hexarthrius rhinoceros ssp. rhinoceros (OLIVIER); Mizunuma & Nagai, 1994:26, fig.
76(a-b), p. 258, pl. 83, 139.
Java : W. Java : Mt. Halimun.
Java : W. Java : Mt. Gede.
Java : E. Java : Mt. Kawi.
Java : Wangoen-Tjisampara.
Hexarthrius rhinoceros ssp. chaudoiri DEYROLLE, 1864.
Hexarthrius chaudoiri DEYROLLE, 1864 [Sumatra].
Hexarthrius rhinoceros ssp. chaudoiri DEYROLLE; Mizunuma & Nagai, 1994:26, 28, fig.
77(a-b), p. 258, pl. 83, 139.
Sumatra (TYPUS).
Sumatra : W. Sumatra Prov. : Padang.
Sumatra : W. Sumatra Prov. : Bukittingi.
Hexarthrius rhinoceros ssp. sadaoi MIZUNUMA, 1994.
Hexarthrius rhinoceros ssp. sadaoi MIZUNUMA in MIZUNUMA & NAGAI, 1994:26, 28, fig.
78(a-b), pl. 139, fig. 343(8-9) [Pagai I.].
Sumatra : W. Sumatra Prov. : Mentawei Is. :
Siberut I., IV-1993 (2 males PARATYPUS in col. T. Mizunuma, Japan).
Sumatra : W. Sumatra Prov. : Mentawei Is. :
Sipora I.
Sumatra : W. Sumatra Prov. : Mentawei Is. :
Pagai I., II-1993 (male HOLOTYPUS in Kanagawa Prefectural Museum, Japan) (3
males PARATYPUS in col. T. Mizunuma, Japan).
Bibliography.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Hexarthrius vitalisi DIDIER, 1925.
Didier, 1925:202; Didier & Séguy,
1953:89; Benesh, 1960:130; Bomans, 1991:146; Maes, 1992:22; Kawano, 1993:12;
Kawano, 1994:9; Mizunuma & Nagai, 1994:259, pl. 85; Kawano, 1997:454.
Thailand
: Chiang Rai (Wiangpapao), Chiang Mai (Chiang Mai--18°50N-98°57E), Lampang
(Muang Pan);
Vietnam :
Tonkin (Tam Dao near Vinh Yen, Paklay*, Chapa*).