Sphaenognathus (Sphaenognathus) BUQUET, 1838

(G. = Sphaenognathus prionoides)

= Orthognathus DEJEAN 1837 (G. = Orthognathus prionoides)

= Sphoenognathus BUQUET 1839

= Sphenognathus REICHE 1852


 

Sphaenognathus (Sphaenognathus) albofuscus (BLANCHARD, 1846).

Orthognathus albofuscus BLANCHARD, 1846:193, pl. 12 [Peru] [Sphenognathus].

Burmeister, 1847:341; Parry, 1870:69; Blackwelder, 1944:195; Blackwelder, 1982:195; Adam, 1986:15; Chalumeau & Brochier, 1993:22

Sphaenognathus albofuscus BLANCHARD; Mizunuma & Nagai, 1994:207, 209, pl. 5, fig. 55(1).

Colombia (1 male citated by Mizunuma & Nagai 1994).

? Venezuela

? Ecuador

? Peru (TYPUS of Orthognathus albofuscus BLANCHARD).

Bolivia

BIBLIOGRAPHY

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Sphaenognathus (Sphaenognathus) alticollis (MOLLENKAMP, 1912).

Sphenognathus alticollis MOLLENKAMP, 1912:302 [Peru].

Blackwelder, 1944:195; Benesh, 1960:149; Lacroix, 1990:12; Blackwelder, 1982:195

Sphaenognathus matheusi BARTOLOZZI, BOMANS & ONORE, 1992:211 [Ecuador : Napo : La Bonita].

Maes, 1992:32; Chalumeau & Brochier, 1993:22

Sphaenognathus alticollis MOLLENKAMP; Mizunuma & Nagai, 1994:208, pl. 5, fig. 47(1-2).

Ecuador : Napo : La Bonita

Peru (TYPUS of Sphenognathus alticollis MOLLENKAMP).

Peru : San Martín : Río Huallaga, I-1985 (2 males citated by Mizunuma & Nagai 1994).

BIBLIOGRAPHY

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Sphaenognathus (Sphaenognathus) armatus (PARRY, 1872).

Sphenognathus armatus PARRY, 1872:73, pl. 1 [Colombia].

Blackwelder, 1944:195; Blackwelder, 1957:1397; Benesh, 1960:149; Blackwelder, 1982:195, 1397; Lacroix, 1987:12; Maes, 1992:32; Chalumeau & Brochier, 1993:22.

Sphenognathus armatus PARRY; Mizunuma & Nagai, 1994:208.

Colombia (TYPUS of Sphenognathus armatus PARRY).

Colombia : Bogota.

BIBLIOGRAPHY

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Sphaenognathus (Sphaenognathus) bellicosus (BOILEAU, 1904).

Sphenognathus bellicosus BOILEAU, 1904:277, 284 [Sierra Nevada de Santa Marta].

Boileau, 1905:38; Blackwelder, 1944:195; Benesh, 1960:149

Sphaenognathus nobilis ssp. colombiensis MOXEY, 1962:199, 201-202 [Colombia : Santa Marta].

Sphaenognathus rehni MOXEY, 1962:199, 201-202 [Colombia : Santa Marta].

Lacroix, 1969:591, 594; Howden & Campbell, 1974:109-114; Blackwelder, 1982:195; Maes, 1992:32, 34; Bartolozzi & Onore, 1993:127; Chalumeau & Brochier, 1993:22

Sphaenognathus bellicosus BOILEAU; Mizunuma & Nagai, 1994:207, pl. 4, fig. 43(1-3).

Colombia : Magdalena : Sierra Nevada de Santa Marta, 11°N-75°30W (TYPUS of Sphenognathus bellicosus BOILEAU).

Colombia : Magdalena : Sierra Nevada de Santa Marta, 11°N-75°30W : San Lorenzo, 8-V-1973 (1 male citated by Mizunuma & Nagai 1994).

Colombia : El Libano (1 male citated by Mizunuma & Nagai 1994).

Colombia, 1981 (1 female citated by Mizunuma & Nagai 1994).

BIBLIOGRAPHY

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Sphaenognathus (Sphaenognathus) curvipes (BENESH, 1948) [Sphoenognathus, Sphenognathus].

Benesh, 1948:45, pl.1; Benesh, 1960:150; Lacroix, 1972:38, 40; Lacroix, 1987:12; Maes, 1992:32; Chalumeau & Brochier, 1993:16; Chalumeau & Brochier, 1993:22.

Bolivia : Incachaca*.

 

Sphaenognathus (Sphaenognathus) feisthamelii ssp. feisthamelii (GUERIN-MENEVILLE, 1838).

Chiasognathus feisthamelii GUERIN-MENEVILLE, 1838:287 [Nova Granada] [Sphenognathus] [festhameli].

Guerin-Meneville, 1839:pl.39; Burmeister, 1847:340; Murray, 1857

Sphenognathus circumflexus PARRY, 1874:367, 368, pl. 4 [Sphaenognathus] [Venezuela].

Sphenognathus canaliculatus PARRY, 1874:367, 368, pl. 4 [Sphaenognathus] [Venezuela].

Boileau, 1911:428, pl.33; Boileau, 1913:215; Kriesche, 1922:115; Nagel, 1924:396; Blackwelder, 1944:195; Didier & Séguy, 1953:24, 72; Benesh, 1960:150

Sphenognathus boileaui LACROIX, 1969:587, 589, 591, 594 [Sphaenognathus] (in part) [Peru : Huanaco].

Sphenognathus villiersi LACROIX, 1969:587, 589, 591, 594 [Sphaenognathus] [Colombia : Iconongo].

De Lisle, 1970:93; Lacroix, 1972:50

Sphenognathus inflexus DE LISLE, 1975:262-264.

Blackwelder, 1982:195; Lacroix, 1985:5; Adam, 1986:15, 17

Sphaenognathus subtilis LACROIX, 1987:13 [Ecuador : Napo : Cordillera de Huacamayos].

Lacroix, 1988:5; Lacroix, 1989:5; Bartolozzi, Bomans & Onore, 1992:195, 199, 226; Maes, 1992:32, 34; Bartolozzi, Onore & Bomans, 1993:169-172; Chalumeau & Brochier, 1993:22; Onore, 1994:94

Sphaenognathus feisthameli (GUERIN-MENEVILLE); Mizunuma & Nagai, 1994:207, pl. 4, fig. 44(1-3).

Sphaenognathus canaliculatus PARRY; Mizunuma & Nagai, 1994:208.

Sphaenognathus circumflexus PARRY; Mizunuma & Nagai, 1994:208.

Bartolozzi, 1996:487.

Colombia = Nova Granada (TYPUS of Chiasognathus feisthamelii GUERIN-MENEVILLE).

Colombia : Bogota

Colombia : Iconongo

Venezuela : Merida : El Vigia

Ecuador : Napo : Cordillera de Huacamayos

Ecuador : Napo : Cosanga

Ecuador : Napo : Loma Playa : Cosanga

Ecuador : Loja : Loja

Peru : Huánuco : Huánuco

Peru : Oxapampa, II/IV-1984 (2 males, 1 female citated in Mizunuma & Nagai 1994).

? Bolivia.

 

Sphaenognathus (Sphaenognathus) feisthamelii ssp. giganteus (BOILEAU 1911).

Sphenognathus giganteus BOILEAU, 1911 [Chulumani].

Sphenognathus boileaui LACROIX, 1969 [Sphaenognathus] (in part) [Peru : Huanaco].

NOTE : Chalumeau & Brochier (1993) considers S. giganteus as a good species.

Peru : Huanaco

Bolivia : Cochabamba : Cochabamba

Bolivia : Conreo

Bolivia : Chulumani (TYPUS of Sphenognathus giganteus BOILEAU).

BIBLIOGRAPHY

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Sphaenognathus furumii LACROIX, 1990.

Sphaenognathus furumii LACROIX, 1990:11-12, pl. 1 [Yungas].

Chalumeau & Brochier, 1993:22

Sphaenognathus furumii LACROIX; Mizunuma & Nagai, 1994:208, pl. 5, fig. 46.

NOTE : Chalumeau & Brochier considers that the Holotype of this species is synonym of S. alticollis, but that the paratypes are possibly a good species. So if the name S. furumii is a synonym of S. alticollis, they propose S. cameneni as a new name for the remains of paratypes of S. furumii.

Bolivia : La Paz : Yungas (TYPUS of Sphaenognathus furumii LACROIX).

Bolivia : La Paz : Yungas, 21-XI-1984 (2 males citated by Mizunuma & Nagai 1994).

Bolivia : La Paz : Yungas, XII-1984 (1 female citated by Mizunuma & Nagai 1994).

BIBLIOGRAPHY

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Sphaenognathus (Sphaenognathus) garleppi (BOILEAU, 1899) [Sphenognathus].

= Sphenognathus manifestus JAKOWLEW 1901 [Bolivia not Venezuela].

= Sphenognathus inca KRIESCHE 1922 [Peru : Chanchamayo].

= Sphaenognathus cartereti LACROIX 1987 [Ecuador : Río de Baños].

Boileau, 1899:196; Jakowlew, 1901:176; Boileau, 1913:215; Kriesche, 1922:115-116; Blackwelder, 1944:195; Didier & Séguy, 1953:24; Blackwelder, 1957:1397; Benesh, 1960:150; Lacroix, 1969:591, 594; De Lisle, 1975:262; Blackwelder, 1982:195, 1397; Klausnitzer, 1982:26; Adam, 1986:15; Lacroix, 1987:11; Bartolozzi, Bomans & Onore, 1992:192; Maes, 1992:32, 33; Chalumeau & Brochier, 1993:22.

Venezuela;

Ecuador : Tungurahua (Río de Baños);

Peru : Chanchamayo;

Bolivia* : La Paz (Youngas), Santa Cruz (Santa Cruz), Yucigase, Tanampaya.

 

Sphaenognathus (Sphaenognathus) higginsi (PARRY, 1876) [Chiasognathus, Sphenognathus].

Parry, 1876:174; Boileau, 1913:215; Blackwelder, 1944:195; Benesh, 1960:150; Benesh, 1960:150; Lacroix, 1972:47; Blackwelder, 1982:195; Bartolozzi, Bomans & Onore, 1992:238; Maes, 1992:33; Chalumeau & Brochier, 1993:22.

? Ecuador;

Bolivia*.

 

Sphaenognathus (Sphaenognathus) lindenii MURRAY, 1857.

Sphaenognathus lindenii MURRAY, 1857:221, pl. 3 [Quito] [Sphenognathus].

Sphenognathus murrayi THOMSON, 1862:407, 409 [Sphaenognathus] [Venezuela].

Sphaenognathus rugosum PARRY, 1863:445 (nomen nudum).

Sphaenognathus kolbei KRIESCHE, 1919 (Syntypus female) [Ecuador : Gualea].

Blackwelder, 1944:195; Benesh, 1948:pl.1; Benesh, 1960:150; Lacroix, 1972:37, 44; De Lisle, 1967:522; Lacroix, 1972:37; De Lisle, 1974:784-785; Blackwelder, 1982:195; Adam, 1986:15, 17; Lacroix, 1987:12; Bartolozzi, Bomans & Onore, 1992:204; Maes, 1992:32-33; Bartolozzi, Onore & Bomans, 1993:168; Chalumeau & Brochier, 1993:16; Chalumeau & Brochier, 1993:22; Onore, 1994:88-92

Sphaenognathus lindenii MURRAY; Mizunuma & Nagai, 1994:209, pl. 5, fig. 56(1-2).

? Venezuela : Merida

Ecuador : Pichincha : Quito (TYPUS of Sphaenognathus lindenii MURRAY).

Ecuador : Pichincha : La Cocha, 2000 m, III-1989 (1 male, 1 female citated in Mizunuma & Nagai 1994).

Ecuador : Pichincha : Aloag

Ecuador : Pichincha : Gualea

? Peru

Subterranean larvae, in humus-rich soil.

BIBLIOGRAPHY

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Sphaenognathus (Sphaenognathus) metallescens (DE LISLE, 1967) [Sphenognathus].

De Lisle, 1967:521; Maes, 1992:33; Bartolozzi, Onore & Bomans, 1993:165-169; Chalumeau & Brochier, 1993:22.

Colombia : Popayan*, Cauca--Silvia.

 

Sphaenognathus (Sphaenognathus) metallifer BOMANS & LACROIX, 1972.

Sphaenognathus metallifer BOMANS & LACROIX en LACROIX, 1972:42 [El Tambo] [Sphenognathus].

Sphenognathus oberon KRIESCHE; De Lisle, 1974:785 (female) [Sphaenognathus] [Ecuador : El Tambo].

Bartolozzi, Bomans & Onore, 1992:214; Maes, 1992:33; Chalumeau & Brochier, 1993:22; Onore, 1994:92

Sphaenognathus metallifer BOMANS & LACROIX; Mizunuma & Nagai, 1994:209, pl. 5, fig. 57(1).

Ecuador : Cañar : El Tambo (TYPUS of Sphaenognathus metallifer BOMANS & LACROIX).

Ecuador : Cañar : El Tambo, 2800 m, 5-III-1965 (1 male citated by Mizunuma & Nagai 1994).

Ecuador : Cañar : 11 mi. N. Juncal

BIBLIOGRAPHY

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Sphaenognathus (Sphaenognathus) nobilis (PARRY, 1874).

Sphenognathus nobilis PARRY, 1874:366, pl. 4 [Venezuela].

Zang, 1905:211; Blackwelder, 1944:195; Benesh, 1960:150; Lacroix, 1969:591, 594; De Lisle, 1970:93; Blackwelder, 1982:195; Onore & Bordon, 1990(1992):97-98; Maes, 1992:33; Chalumeau & Brochier, 1993:16; Chalumeau & Brochier, 1993:22; Onore, 1994:92-93

Sphaenognathus nobilis ssp. nobilis PARRY; Mizunuma & Nagai, 1994:207, pl. 4, fig. 45(1).

Venezuela (TYPUS of Sphenognathus nobilis PARRY).

Venezuela : Caracas

Venezuela : Escorial

Venezuela : Merida : Sierra Nevada National Park : La Mucuy

Venezuela (1 male citated by Mizunuma & Nagai 1994).

Subterranean larvae, in humus-rich soil.

BIBLIOGRAPHY

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Sphaenognathus (Sphaenognathus) oberon (KRIESCHE, 1922).

Sphenognathus oberon KRIESCHE, 1922:116 [Telimbela].

Blackwelder, 1944:195; Benesh, 1960:150; Lacroix, 1972:43; De Lisle, 1974:785; Blackwelder, 1982:195; Bartolozzi & Onore, 1992; Bartolozzi, Bomans & Onore, 1992:219; Maes, 1992:33; Bartolozzi & Onore, 1993:126-128; Chalumeau & Brochier, 1993:22; Onore, 1994:93

Sphaenognathus oberon KRIESCHE; Mizunuma & Nagai, 1994:209, pl. 5, fig. 58(1-2).

Ecuador : Bolivar : Telimbela (TYPUS of Sphenognathus oberon KRIESCHE).

Ecuador : Bolivar : Totoras, III-1992 (1 male, 1 female citated by Mizunuma & Nagai 1994).

Ecuador : Bolivar : Mt. Polvora

Ecuador : Bolivar : trail to Porvenir : Hacienda Talahua

Subterranean larvae, in humus-rich soil.

BIBLIOGRAPHY

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Sphaenognathus (Sphaenognathus) prionoides (BUQUET, 1838).

Dejean, 1837:193

Orthognathus prionoides BUQUET, 1838:104 [Colombia] [Sphoenognathus, Sphenognathus].

Buquet, 1839:pl.1; Burmeister, 1847:341; Heyne & Taschenberg, 1908:48

Sphaenognathus prionoides var. bispinosus DIDIER, 1928:11 (nomen nudum).

Sphaenognathus prionoides var. spinosus DIDIER, 1928:11 (nomen nudum).

Blackwelder, 1944:195; Didier & Séguy, 1953:24; Benesh, 1960:150-151

Sphenognathus pristis DE LISLE, 1970:91 [Peru : Cumbase].

Sphenognathus dentifer LACROIX, 1972:33, 34, 38 [Colombia].

Moore, 1978:99; Blackwelder, 1982:195; Klausnitzer, 1982:44

Sphaenognathus bidentatus ADAM, 1986:15 [Chile].

Lacroix, 1987:12; Maes, 1992:32, 33; Chalumeau & Brochier, 1993:22

Sphaenognathus pristis DE LISLE; Mizunuma & Nagai, 1994:208.

Sphaenognathus dentifer LACROIX; Mizunuma & Nagai, 1994:208.

Colombia : Bogota

Colombia : Muzo;

Colombia (TYPUS of Orthognathus prionoides BUQUET).

Venezuela : Merida

Ecuador : Loja : Loja

Peru : Cumbase

Bolivia

? Chile : La Araucanía : Victoria

BIBLIOGRAPHY

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Sphaenognathus (Sphaenognathus) pubescens (WATERHOUSE, 1873) [Chiasognathus, Sphenognathus].

= Sphaenognathus villosus ADAM 1986 [Colombia*].

Waterhouse, 1873:110; Parry, 1874:pl.5; Heyne & Taschenberg, 1908:48; Blackwelder, 1944:195; Benesh, 1960:151; De Lisle, 1970:93; Blackwelder, 1982:195; Adam, 1986:16; Lacroix, 1987:12; Maes, 1992:33, 34; Chalumeau & Brochier, 1993:24.

Colombia;

Venezuela*.

 

Sphaenognathus (Sphaenognathus) rotundatus LACROIX, 1987.

Sphaenognathus rotundatus LACROIX, 1987:12 [Cordillera de Merida].

Lacroix, 1987:12; Maes, 1992:34; Chalumeau & Brochier, 1993:24

Sphaenognathus rotundatus LACROIX; Mizunuma & Nagai, 1994:208.

NOTE : For Chalumeau & Brochier (1993), this species could be a synonym of S. taschenbergi.

Venezuela : Cordillera de Merida (TYPUS of Sphaenognathus rotundatus LACROIX).

BIBLIOGRAPHY

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Sphaenognathus (Sphaenognathus) signatus (PARRY, 1874).

Sphenognathus signatus PARRY, 1874:366, pl. 5 [Venezuela].

Sphenognathus praestans JAKOWLEW, 1901:77 [Sphaenognathus] [Venezuela].

Blackwelder, 1944:195; Benesh, 1960:150, 151

Sphenognathus hemisphanestus DE LISLE, 1975:264 [Sphaenognathus] [Colombia : Paso del Aunindin].

Blackwelder, 1982:195; Lacroix, 1990:12; Maes, 1992:33, 34; Chalumeau & Brochier, 1993:24

Sphaenognathus signatus PARRY; Mizunuma & Nagai, 1994:208.

Sphaenognathus hemiphanestus DE LISLE; Mizunuma & Nagai, 1994:208.

Colombia : Paso del Aunindin

? Venezuela (TYPUS of Sphenognathus signatus PARRY).

BIBLIOGRAPHY

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Sphaenognathus (Sphaenognathus) spinifer (BOILEAU, 1904).

Sphenognatus spinifer BOILEAU, 1904:284 [Colombia].

Blackwelder, 1944:195; Benesh, 1960:151; Lacroix, 1972:37; Blackwelder, 1982:195

Sphenognathus lachaumei LACROIX, 1983:5 [Colombia : Bianchi].

Lacroix, 1990:12; Téllez, 1990:38; Maes, 1992:33, 34; Chalumeau & Brochier, 1993:24

Sphaenognatus spinifer BOILEAU; Mizunuma & Nagai, 1994:208.

Sphaenognathus lachaumei LACROIX; Mizunuma & Nagai, 1994:208.

Colombia (TYPUS of Sphaenognatus spinifer BOILEAU).

Colombia : Bianchi

Colombia : Cauca : Pereira

BIBLIOGRAPHY

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Sphaenognathus (Sphaenognathus) taschenbergi (PARRY, 1874).

Sphenognathus taschenbergi PARRY, 1874:368, pl. 5 [Venezuela] [tachenbergi].

Blackwelder, 1944:195; Benesh, 1960:151

Sphenognathus didieri LACROIX, 1972:37, 38, 40, 44 [Venezuela : Cord. Merida--El Vigia].

Blackwelder, 1982:195; Klausnitzer, 1982:44; Lacroix, 1987:12; Maes, 1992:34

? Sphaenognathus taschenbergi var. paramoensis CHALUMEAU & BROCHIER, 1993:24.

NOTE : Chalumeau & Brochier (1993) wrote that var. paramoensis could be a twin species with S. taschenbergi.

Sphaenognathus taschenbergi PARRY; Mizunuma & Nagai, 1994:208.

Sphaenognathus didieri LACROIX; Mizunuma & Nagai, 1994:208.

Sphaenognathus taschenbergi PARRY; Chalumeau & Brochier, 2001:595.

Venezuela (TYPUS of Sphenognathus taschenbergi PARRY).

Venezuela : Cordillera de Merida : El Vigia

BIBLIOGRAPHY

CHALUMEAU F. & BROCHIER B. (2001) Une forme fossile nouvelle de Chiasognathinae : Protognathinus spielbergi (Coleoptera,  Lucanidae). Lambillionea, 101(4):593-595.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Sphaenognathus sp.

Sphaenognathus sp. Mizunuma & Nagai, 1994:208, lam. 5, fig. 48(1-2), 49(1), 50 (1), 51(1-2).

Colombia : Buga Valley, II-1985 (2 males citated by Mizunuma & Nagai 1994).

Colombia : Paso del Aunindin, II-1985 (1 male citated by Mizunuma & Nagai 1994).

Colombia, II-1985 (1 female citated by Mizunuma & Nagai 1994).

Colombia (1 male, 1 female citated by Mizunuma & Nagai 1994).

BIBLIOGRAPHY

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 


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