Lucanus (Pseudolucanus) HOPE, 1845
(G. =
Scarabaeus capreolus)
= Eolucanus
KUROSAWA 1970 (G. = Pseudolucanus gracilis)
Lucanus
(Pseudolucanus) atratus HOPE, 1831.
Lucanus
atratus HOPE, 1831:22 [Nepal].
Lucanus
atratus HOPE; Hope & Westwood, 1845:10
Lucanus
atratus HOPE; Burmeister, 1847:528
Lucanus
atratus HOPE; Reiche, 1853:71
Lucanus
atratus HOPE; Planet, 1895:144
Pseudolucanus
oberthuri PLANET, 1896:278 (female) [Sikkim].
Lucanus
atratus HOPE; Planet, 1899:9
Lucanus
atratus HOPE; Planet, 1902:pl.1
Lucanus
atratus HOPE; Heyne & Taschenberg, 1908:50
Lucanus
atratus HOPE; Boileau, 1913:218
Lucanus
atratus HOPE; Gravely, 1915:411
Lucanus
atratus HOPE; Arrow, 1950:58, pl.4
Lucanus atratus HOPE; Didier & Séguy, 1952:pl.9
Lucanus atratus HOPE; Didider & Séguy, 1953:77
Lucanus atratus HOPE; Benesh, 1960:131
Lucanus atratus HOPE; Endrodi, 1968:49
Lucanus atratus HOPE; Kurosawa, 1970
Lucanus atratus HOPE; Endrodi, 1971:10
Lucanus atratus HOPE; Nakane, 1972:41
Lucanus atratus HOPE; De Lisle, 1975:204-205
Lucanus atratus HOPE; Maes, 1992:20
Lucanus
atratus HOPE; Bomans, 1994:493
Lucanus
atratus HOPE; Bartolozzi & Sforzi, 1994:113, 114
Lucanus
atratus HOPE; Boucher, 1994:506-507
Lucanus
atratus HOPE; Mizunuma & Nagai, 1994:211, pl. 7, fig. 78(1-7).
Lucanus
atratus HOPE; Schenk, 1996:510
Lucanus
atratus HOPE; Araya, 1996:16.
Lucanus
atratus HOPE & WESTWOOD; Araya, 2001:10-11, figs. 6-7.
·
India : Sikkim
·
India : West Bengal : Darjeeling : Tiger Hill, VI-1986 (3 males citated
by Mizunuma & Nagai 1994).
·
India : West Bengal : Ghoom
·
India : West Bengal : Kurseong
·
India :?Assam
·
Nepal, col. Hope (male TYPUS of Lucanus atratus HOPE, 1831, #349
in Oxford University Museum of Natural History).
·
Nepal : Thodung
·
Nepal : Gosainkund
·
Nepal : Godavari : Mt. Phulcoki, VI-1990 (2 males, 1 female citated by
Mizunuma & Nagai 1994).
·
Nepal : Mt. Phulcoki : Kathmandou
·
Nepal : Myagdi Distr. : Dhawalagiri : Ghorepani
·
Nepal : Helambu : Sundarijal - Kutumsang
·
Nepal : Bagmati : Nuwakot : Gul Bhanjyang
·
Nepal : Mure
·
Nepal : Rupchet Ganesh
·
Nepal : Arun Valley
·
Nepal : Sapt Kosi
·
Nepal : Dhaulagiri
·
Nepal : Gunsa
·
Nepal : Chitrei
·
Nepal, VII-1973 (1 male citated by Mizunuma & Nagai 1994).
|
|
|
Male TYPE of of Lucanus atratus HOPE,
1831, from Nepal (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from
Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Lucanus (Pseudolucanus) barbarossa FABRICIUS, 1801.
Lucanus
barbarossa FABRICIUS, 1801:251 or 261 [Morocco : Tanger].
Lucanus
barbarossa FABRICIUS; Illiger, 1803:233
Lucanus
barbarossa FABRICIUS; Illiger, 1805:104
Lucanus
barbarossa FABRICIUS; Schonherr, 1817:325
Lucanus
barbarossa FABRICIUS; Burmeister, 1847:349
Lucanus
barbarossa FABRICIUS; Duval, 1859:13, pl.1
Lucanus
barbarossa FABRICIUS; Kraatz, 1860:60, 265, pl.7
Lucanus
barbarossa FABRICIUS; Martinez & Saez, 1873.
Lucanus
barbarossa FABRICIUS; Reitter, 1891:141
Lucanus barbarossa FABRICIUS; Paulino de Oliveira, 1894.
Lucanus barbarossa FABRICIUS; Planet,1895:180
Lucanus barbarossa FABRICIUS; Correa de Barros, 1896.
Lucanus barbarossa FABRICIUS; Planet, 1899:26, pl.4
Lucanus barbarossa FABRICIUS; Scabra, 1905:8, pl.
Lucanus barbarossa FABRICIUS; Barthe, 1909:6
Lucanus barbarossa FABRICIUS; Martinez de la Escalera, 1914:1-553
Lucanus barbarossa FABRICIUS; Fuente, 1926:21-36
Lucanus barbarossa FABRICIUS; Seabra, 1943.
Lucanus barbarossa FABRICIUS; Cobos, 1949:563-609
Lucanus barbarossa FABRICIUS; Didier, 1949:pl.8
Lucanus barbarossa FABRICIUS; Ladeiro, 1950.
Lucanus barbarossa FABRICIUS; Kocher, 1958:1-83
Lucanus barbarossa FABRICIUS; Benesh, 1960:131
Lucanus barbarossa FABRICIUS; Carrion, 1961:99-126
Lucanus barbarossa FABRICIUS; Baguena, 1967:576 pp.
Lucanus barbarossa FABRICIUS; Español, 1973:99-111
Lucanus barbarossa FABRICIUS; Español & Belles, 1982:73
Lucanus barbarossa FABRICIUS; García-Parrón & Benitez-Donoso,
1984:79-82
Lucanus barbarossa FABRICIUS; Andujar-Tomas, Lencina-Gutierrez &
Ruano-Marco, 1988:61-65
Lucanus barbarossa FABRICIUS; Lopez-Colon, 1992:89-90
Lucanus barbarossa FABRICIUS; Maes, 1992:20
Lucanus barbarossa FABRICIUS; Baraud, 1993:42-64
Lucanus barbarossa FABRICIUS; López Vaamonde, Pino Pérez, Devesa
Regueiro & Martínez Fernández, 1993:349
Lucanus barbarossa FABRICIUS; Mizunuma & Nagai, 1994:212, pl.7
Lucanus barbarossa FABRICIUS; Ruiz & Avila, 1995:208-209
Pseudolucanus barbarossa (FABRICIUS); López-Colón, González Peña & Beltrán
Valen, 1996:16, 17, 18.
Lucanus barbarossa FABRICIUS; Kawano, 1997:454.
Pseudolucanus barbarossa (FABRICIUS); Bahillo de la Puebla, 2001:165.
·
Portugal : Arila
·
Spain : Cantabria :
Santander, 43°28 N - 3°48 W
·
Spain : Orense : Supra
Vilar de Silva : Rubiá
·
Spain : Huesca : Santa
Cilia de Jaca, 42°34 N - 0°43 W (citated by López-Colón, González Peña &
Beltrán Valen, 1996).
·
Spain : Huesca :
Agüero, 42°22 N - 0°54 W (citated by López-Colón, González Peña & Beltrán
Valen, 1996).
·
Spain : Huesca :
Loarre, 42°19 N - 0°37 W (citated by López-Colón, González Peña & Beltrán
Valen, 1996).
·
Spain : Huesca : Graus,
42°11 N - 0°20 W (citated by López-Colón, González Peña & Beltrán Valen,
1996).
·
Spain : Huesca : La
Peña (citated by López-Colón, González Peña & Beltrán Valen, 1996).
·
Spain : La Rioja :
Logroño, 42°28 N - 2°27 W
·
Spain : Castilla – León
: Burgos, 42°20 N – 3°42 W
·
Spain : León : Castilla
La Mancha : Matallana, 41°03 N - 3°20 W
·
Spain : Alava
·
Spain : Navarra :
Monasterio de Irache, 5-VII-1999 (citado por Bahillo de la Puebla, 2001).
·
Spain : Zaragoza :
Luesia, 42°22 N - 1°01 W (citated by López-Colón, González Peña & Beltrán
Valen, 1996).
·
Spain : Zaragoza : El
Buste, 41°53 N - 1°36 W (citated by López-Colón, González Peña & Beltrán
Valen, 1996).
·
Spain : Zaragoza : Litago,
41°49 N - 1°45 W (citated by López-Colón, González Peña & Beltrán Valen,
1996).
·
Spain : Zaragoza : Vera
de Moncayo, 41°47 N - 1°50 W (citated by López-Colón, González Peña &
Beltrán Valen, 1996).
·
Spain : Zaragoza :
Cosuenda, 41°22 N - 1°18 W (citated by López-Colón, González Peña & Beltrán
Valen, 1996).
·
Spain : Zaragoza :
Agramonte (citated by López-Colón, González Peña & Beltrán Valen, 1996).
·
Spain : Zaragoza : Biel
(citated by López-Colón, González Peña & Beltrán Valen, 1996).
·
Spain : Valladolid, 41°39
N – 4°43 W
·
Spain : Lérida, 41°37 N
– 0°37 E
·
Spain : Barcelona,
41°23 N – 2°11 E
·
Spain : Vieja Castilla
: Barruecopardo, 41°04 N - 6°40 W
·
Spain : Salamanca,
40°58 N – 5°39 W
·
Spain : Guadalajara,
40°38 N – 3°10 W
·
Spain : Avila : Las
Navas de Marques, 40°37 N – 4°20 W
·
Spain : Teruel :
Orihuela de Tremedal, 40°33 N - 1°39 W (citated by López-Colón, González Peña
& Beltrán Valen, 1996).
·
Spain : Teruel :
Rubielos de Mora, 40°12 N - 0°39 W (citated by López-Colón, González Peña &
Beltrán Valen, 1996).
·
Spain : Madrid, 40°26 N – 3°41 W
·
Spain : Castellón
·
Spain : Cuenca, 40°05 N
– 2°08 W
·
Spain : Cáceres, 39°28 N – 6°22 W
·
Spain : Toledo, 39°52 N – 4°02 W
·
Spain : Valencia, 39°29 N – 0°21 W
·
Spain : Albacete, 39°00 N – 1°52 W
·
Spain : Ciudad Real,
38°59 N – 3°56 W
·
Spain : Granada, 37°11
N – 3°36 W
·
Spain : Almería, 36°51
N – 2°28 W
·
Spain : Cádiz : Sierra
de Ronda, 36°45 N - 5°10 W : Nava de San Luis
·
Spain : Cádiz : Sierra
de Ronda, 36°45 N - 5°10 W : Puerto de los Pilones
·
Spain : Málaga :
Antequera, 37°01 N - 4°33 W
·
Spain : Cazurro : Los
Jurdes
·
Morocco : Tanger, 35°47 N - 5°48 W (TYPUS of Lucanus barbarossa FABRICIUS,
1801).
·
Morocco : High Atlas
·
Morocco : Middle Atlas : Ifrane, 33°32 N - 5°07 W
·
Morocco : Middle Atlas : Azrou, 33°27 N - 5°14 W
·
Morocco : Leiria
·
Morocco : Marrakesh, 31°38 N - 8°00 W
·
Morocco : Rif : Talassentane
·
Algeria
·
Tunisia
|
|
|
|
Pseudolucanus barbarossa (España, fotos de un
macho tomadas por Ignasi Serrahima) |
|
|
|
|
|
Pseudolucanus barbarossa (España, foto de un macho
tomada por Miguel Ángel Pinto). |
|
BIBLIOGRAFIA
BAHILLO DE LA
PUEBLA P. (2001)
El Camino de Santiago, una excursión entomológica (Andanzas de un escarabajero
por la ruta jacobae). Bol. S.E.A., España, 28:157-165.
BARAUD J. (1993) Les Coléoptères Lucanoidea de l'Europe et du Nord de l'Afrique. Bull. mens. Soc. linn. Lyon, 62 (2):42-64.
FUENTE J.M. de la (1926) Catálogo sistemático-geografico de los Coleópteros de
la Península Ibérica y Baleares. Bol. Soc. Ent. Espñ., 9(1-2):143-160.
KOCHER L. (1958) Catalogue commenté des Coleópteros du Maroc, fase. VIL LameUicomes. Trav. ¡ns. Scient. Chénfien, ser. wol. núm. 16. Rabal: 1-83.
LOPEZ-COLON J.I. (1992) Nuevos datos sobre Lucanidae y Melolonthidae de
Sierra Nevada (Sur de la Península Ibérica) (Col.). Nouv. Revue Ent. (N.S.), 9 (I): 89-90.
LOPEZ-COLON
J.I., GONZALEZ PEÑA C.F. & BELTRAN VALEN J.R. (1996) Familia : Lucanidae. Catalogus de la
Entomofauna Aragonesa, No. 12:15-20.
MARTINEZ DE LA ESCALERA M. (1914) Los Coleópteros de Marruecos. Trav. del Museo
Nac. de Cieñe. Nat. de Madrid, ser. zool., n° 11: 191.
RUIZ J.L. & AVILA J.M. (1995) Nuevas localizaciones de Pseudolucanus
barbarossa (Fabricius, 1801) y Platycerus spinifer Schauffus, 1862 en Andalucía (Coleóptera: Lucanidae).
Bol. Asoc. Esp. Entomología. 19, 3-4: 208.
SEABRA A.F. de (1943) Contribucoes para o inventario da fauna lusitanica.
Insecta, Coleóptera. Mem. Estud. Mus. Zool. Univ. Coimbra, 142: 1-152.
Lucanus (Pseudolucanus) capreolus (LINNAEUS, 1763).
Lucanus
capreolus LINNAEUS, 1763:391.
Lucanus
capreolus LINNAEUS; Linnaeus, 1764:32
Lucanus
capreolus LINNAEUS; Linnaeus, 1767:560
Lucanus capreolus LINNAEUS; De Geer, 1774:336, pl.19
Lucanus dama FABRICIUS, 1775:2 [Virginia].
Lucanus capreolus LINNAEUS; Fabricius, 1787:1
Lucanus capreolus LINNAEUS; Fabricius, 1787:1
Lucanus capreolus LINNAEUS; Olivier, 1789:15-16, pl.2-3
Lucanus capreolus LINNAEUS; Herbst, 1790:302, pl.34
Lucanus capreolus LINNAEUS; Linnaeus (ed. Gmelin), 1790:1589
Lucanus capreolus LINNAEUS; Fabricius, 1792:237
Lucanus capreolus LINNAEUS; Olivier (ed.Illiger), 1800:63
Lucanus capreolus LINNAEUS; Fabricius, 1801:249
Lucanus capreolus LINNAEUS; Olivier (ed. Sturm), 1802:3, pl.3
Lucanus
muticus THUNBERG, 1806:198-199, 205, pl.12.
Lucanus
trigonus THUNBERG, 1806:198-199, 205, pl.12.
Lucanus
capreolus LINNAEUS; Schonherr, 1817:323, 327
Lucanus capreolus LINNAEUS; Dejean, 1837:193
Lucanus capreolus LINNAEUS; Castelnau, 1840:171
Lucanus capreolus LINNAEUS; Burmeister, 1847:355
Lucanus capreolus LINNAEUS; Saunders, 1881:118
Lucanus capreolus LINNAEUS; Fuchs, 1882:50
Lucanus capreolus LINNAEUS; Packard, 1889:226
Lucanus capreolus LINNAEUS; Planet, 1890:23, pl.1
Lucanus capreolus LINNAEUS; Planet, 1895:154
Lucanus capreolus LINNAEUS; Planet, 1899:24, pl.2
Lucanus
capreolus LINNAEUS; Wickham, 1899:22
Lucanus
capreolus LINNAEUS; Planet, 1902:pl.2-3
Lucanus
capreolus LINNAEUS; Hart, 1907:244
Lucanus
capreolus LINNAEUS; Heyne & Taschenberg, 1908:49, pl.6
Lucanus
capreolus LINNAEUS; Easton, 1909:51
Lucanus
capreolus LINNAEUS; Blatchley, 1910:909
Pseudolucanus
capreolus ab. nigricephalus BENESH, 1938:271, 273 [N. Illinois,
Tennessee--Deer Lodge, Tennessee--Nashville, Pennsylvania--Mt. Airy, Massasuchetts--Wilmington--Gipsy
Moth Lab.].
Lucanus
capreolus (LINNAEUS); Williams, 1945.
Lucanus capreolus LINNAEUS; Didier & Séguy, 1952:pl.12
Lucanus capreolus LINNAEUS; Benesh, 1960:131-132
Lucanus capreolus LINNAEUS; Dillon & Dillon, 1961
Lucanus capreolus LINNAEUS; Holloway, 1963:100
Lucanus capreolus LINNAEUS; Ritcher, 1966
Lucanus
capreolus LINNAEUS; Arnett, 1968
Lucanus
capreolus LINNAEUS; Mathieu, 1969:1055
Lucanus
capreolus LINNAEUS; Hatch, 1971:430
Lucanus
capreolus LINNAEUS; Boucher & Huang, 1991:31
Lucanus
capreolus LINNAEUS; Ratcliffe, 1991:265
Lucanus
capreolus LINNAEUS; Maes, 1992:20
Lucanus
capreolus LINNAEUS; Mizunuma & Nagai, 1994:212, pl.7
Lucanus capreolus LINNAEUS; Poole & Gentili, 1996:297.
Lucanus capreolus (LINNAEUS); Reyes-Castillo, 486.
·
Canada : Ontario
·
Canada : Quebec
·
USA : California
·
USA : Idaho :
Pocatello, 42°50N-112°25W
·
USA : Nebraska : Seward
Co. : 7 mi. S. Garland, 40°55N-96°55W
·
USA : Nebraska : Saline Co. : DeWitt, 40°25N-96°50W
·
USA : Nebraska : Gage Co. : Beatrice, 40°15N-96°45W
·
USA : Nebraska :
Lancaster Co. : Lincoln, 40°45N-96°40W
·
USA : Nebraska : Adams
Co., 40°30N-96°35W
·
USA : Nebraska :
Douglas Co. : Omaha, 41°15N-95°55W
·
USA : Nebraska : Cass Co. : Plattsmouth, 41°N-95°50W
·
USA : Nebraska : Jefferson Co.
USA : Nebraska : Richardson Co. : Indian Cave State Park
·
USA : Nebraska : Sarpy Co. : Schramm Park
·
USA : Kansas
·
USA : Texas : Willascy Co. : Lyford, 26°30N-97°45W
·
USA : Iowa : Iowa City, 41°40N-91°30W
·
USA : Missouri : Kansas City, 39°05N-94°30W
·
USA : Michigan :
Washtenaw
·
USA : Wisconsin
·
USA : Illinois : Lake Co. : Libertyville, 42°25N-88°00W
·
USA : Illinois :
Chicago, 41°50N-87°40W
·
USA : Illinois : Dunes
Park
·
USA : Illinois : Olive
Branch
·
USA : Indiana : Miller
·
USA : Indiana :
Stilesville, 39°40N-86°30W
·
USA : Kentucky
·
USA : Tennessee : Nashville, 36°10N-86°45W
·
USA : Tennessee : Comber Co. : Crossville, 35°55N-85°05W
·
USA : Tennessee : Morgan Co. : Deer Lodge
·
USA : Alabama :
Birmingham, 33°25N-86°50W
·
USA : Ohio
·
USA : New York : Rochester, 43°10N-77°40W
·
USA : New York : New York, 40°45N-73°50W
·
USA : Massachussetts : Chicopee, 42°10N-72°40W
·
USA : Massachussetts : Wilmington
·
USA : Connecticut : Putnam Co., 41°50N-40°50W
·
USA : Pennsylvania : Cornwells
·
USA : Pennsylvania : Pittsburgh, 40°30N-80°00W
·
USA : Pennsylvania : Airy Mts.
·
USA : Maryland : Plummers Island
USA : Delaware
·
USA : West Virginia
·
USA : Virginia
·
USA : North Carolina : Union Co. : Waxhaw, 34°55N-80°45W
·
USA : South Carolina :
Columbia, 34°00N-81°10W
Saproxylophagous :
Fagaceae : Quercus, Fagus.
Magnoliaceae : Liriodendron.
Bibliografía :
REYES-CASTILLO P. (2002) 27. Lucanidae. Pp. 485-492. En LLORENTE
BOUSQUETS J. & MORRONE J.J. Biodiversidad, Taxonomía y Biogeografía de Artrópodos
de México : Hacia una síntesis de su conocimiento. Universidad Nacional
Autónoma de México. Vol. III:692 pp.
WILLIAMS J.L. (1945)
The anatomy of internal genitalia of some Coleoptera. Proc. Ent. Soc. Wash.,
47(4):73-91.
Lucanus
(Pseudolucanus) confusus BOUCHER, 1994.
Pseudolucanus
atratus HOPE; Planet, 1896
Pseudolucanus
atratus HOPE; Planet, 1899
Pseudolucanus
atratus HOPE; Gravely, 1915
Pseudolucanus
atratus HOPE; Arrow, 1950
Pseudolucanus
atratus HOPE; Didier & Séguy, 1952: pl.ix
Pseudolucanus atratus HOPE; De Lisle, 1975
Pseudolucanus atratus HOPE; Mizunuma & Nagai, 1994
Pseudolucanus confusus BOUCHER, 1994:507-510 [British Bootan : Padong]
Pseudolucanus confusus BOUCHER; Araya, 1996:16.
Pseudolucanus
confusus BOUCHER; Araya, 2001:11-12, figs. 14-15.
·
India : Sikkim : Gnatong*
·
India : Sikkim : Padantsin to Lingtou*
·
India : West Bengal : Darjeeling*
·
India : West Bengal : Darjeeling : Tiger Hills*
·
India : West Bengal : Darjeeling : Ghoom*
·
Nepal : Arun Valley
·
Nepal : Sapt Kosi
·
Bhutan (British Bootan) : Padong, 1913, col. L. Durel (male HOLOTYPE of Pseudolucanus
confusus BOUCHER, 1994 in Museum National d’Histoire Naturelle, Paris).
·
Bhutan : Doriula
·
Bhutan : Sampa-Kotoka.
|
|
|
Male HOLOTYPE of Pseudolucanus confusus BOUCHER,
1994, from British Bootan : Padong (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Pseudolucanus) davidis (DEYROLLE, 1878).
Pseudolucanus
davidis DEYROLLE in DEYROLLE & FAIRMAIRE, 1878:93, pl. 4 [China : Mou-Pin].
Pseudolucanus
davidis DEYROLLE, 1878; Planet, 1895:145
Pseudolucanus
davidis DEYROLLE, 1878; Planet, 1899:16
Pseudolucanus
davidis DEYROLLE, 1878; Planet, 1902:97, pl.2
Pseudolucanus
davidis DEYROLLE, 1878; Didier & Séguy, 1952:pl.10
Pseudolucanus
davidis DEYROLLE, 1878; Benesh, 1960:140
Pseudolucanus
davidis DEYROLLE, 1878; Kurosawa, 1978:143
Lucanus
davidis DEYROLLE, 1878; Kikuta, 1986:134
Lucanus
davidis DEYROLLE, 1878; Boucher & Huang, 1991:31
Lucanus
davidis DEYROLLE, 1878; Maes, 1992:21
Lucanus
davidis (DEYROLLE); Mizunuma & Nagai, 1994:211, pl. 7, pl. 138, fig.
77(1-2).
Lucanus
davidis DEYROLLE, 1878; Schenk, 1996:510.
Lucanus
davidis DEYROLLE, 1878; Araya, 2001:12-13, figs.21-24.
·
China : Sichuan : Kanding : Hailougou Glacier Park, 21/24-VII-1992, leg.
R. Dunda (1 male, 1 female citated in Mizunuma & Nagai 1994).
·
China : Xizang (= Thibet) : Mou-Pin, col. A. David, 1870 (male TYPUS,
female TYPUS of Pseudolucanus davidis DEYROLLE, 1878 in Museum National
d’Histoire Naturelle, Paris).
·
China : Yunnan
|
|
|
|
Male TYPE of Pseudolucanus davidis DEYROLLE,
1878, from China : Xizang : Mou-Pin (adapted from Araya, 2001). |
Female TYPE of Pseudolucanus davidis DEYROLLE,
1878, from China : Xizang : Mou-Pin (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Lucanus
(Pseudolucanus) denticulus BOUCHER, 1995.
Pseudolucanus
denticulus BOUCHER, 1995:441-443 [Mts. Gaoligong].
Lucanus
denticulus BOUCHER; Araya, 2001:13, figs.27-28.
·
China-Myanmar border : Gaoligong Mts., 2500 m, VII-1995, col. H.
Bousquets (female HOLOTYPE of Pseudolucanus denticulus BOUCHER, 1995 in
Museum National d’Histoire Naturelle, Paris).
|
|
|
Female HOLOTYPE of Pseudolucanus denticulus
BOUCHER, 1995, from Mts. Gaoligong (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Pseudolucanus) deuvei (LACROIX, 1988) [Pseudolucanus].
Pseudolucanus deuvei LACROIX, 1988:7, pl.1 [India : Bhadarwar].
Pseudolucanus deuvei LACROIX; Maes, 1992:21
Pseudolucanus deuvei LACROIX; Schenk, 1996:508-512.
·
India : Jammu &
Kashmir : Bhadarwar (TYPUS de Pseudolucanus deuvei LACROIX, 1988).
Lucanus (Pseudolucanus) gracilis ALBERS, 1889.
Lucanus
gracilis ALBERS, 1889:319 [Sikkim].
Lucanus
gracilis ALBERS; Planet, 1902:130
Lucanus
gracilis ALBERS; Arrow, 1935:106
Lucanus
gracilis ALBERS; Arrow, 1950:61, pl.5
Lucanus
gracilis ALBERS; Benesh, 1960:141
Lucanus
gracilis ALBERS; Endrodi, 1968:50
Lucanus
gracilis ALBERS; Kurosawa, 1970
Lucanus
gracilis ALBERS; De Lisle, 1975:204
Lucanus
gracilis ALBERS; Kurosawa, 1978:143
Lucanus
gracilis ALBERS; Kikuta, 1986:134-135
Lucanus
gracilis ALBERS; Sakaino, 1987:17
Lucanus
gracilis ALBERS; Kawano, 1988:26
Lucanus
gracilis ALBERS; Boucher & Huang, 1991:31
Lucanus
gracilis ALBERS; Maes, 1992:21
Lucanus
gracilis ALBERS; Bartolozzi & Sforzi, 1994:114
Lucanus
gracilis ALBERS; Mizunuma & Nagai, 1994:211, pl. 7, fig. 75(1-3).
·
India : Sikkim (TYPUS of Lucanus gracilis ALBERS).
·
India : Sikkim : Ratong Valley
·
India : Sikkim : Mt. Kanchenjunga, 25-VI-1990 (1 male, 2 females citated
by Mizunuma & Nagai 1994).
·
India : Assam
·
Nepal : Thodung
·
Nepal : Khumbu
·
Nepal : Likhu Khola-Tal
·
Nepal : Khumjung
·
Nepal : Dudh Kosi-Tal
·
Nepal : Thangpoche
·
Nepal : Barahbise
·
Bhutan : Gogona
·
Bhutan : Thimphu
·
China : Xizang = Thibet : Rongshar Valley
BIBLIOGRAPHY
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic
Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.
Lucanus
(Pseudolucanus) groulti (PLANET, 1897).
Pseudolucanus
groulti PLANET, 1897:227 [India].
Pseudolucanus
groulti PLANET; Planet, 1899:100
Pseudolucanus
groulti PLANET; Arrow, 1950:55
Pseudolucanus
groulti PLANET; Didider & Séguy, 1953:78
Pseudolucanus
groulti PLANET; Benesh, 1960:141
Pseudolucanus
wittmeri LACROIX, 1983 [in part, Paratypes from Nepal : Kalapani].
Pseudolucanus
groulti PLANET; Maes, 1992:21
Pseudolucanus
groulti PLANET; Bartolozzi & Sforzi, 1994:113, 114, 115
Lucanus
groulti (PLANET); Mizunuma & Nagai, 1994:212, pl. 7, fig. 79(1-2).
Pseudolucanus
groulti PLANET; Schenk, 1996:510-512.
Lucanus
groulti (PLANET); Araya, 2001:15, figs. 40-44.
·
Pakistan : Northwest Frontier
·
India, ex coll. Mniszech (male TYPUS, female TYPE of Pseudolucanus
groulti PLANET, 1897 in Museum National d’Histoire Naturelle, Paris).
·
India : Uttar Pradesh (= United Provinces) : Kunayini
·
India : Uttar Pradesh (= United Provinces) : Bhawali
·
India : NW India (2 males citated by Mizunuma & Nagai 1994).
·
Nepal : Kaski Distr. : Todapani : Ghandrung - Ghorapani
·
Nepal : Kalapani (PARATYPUS of Pseudolucanus wittmeri LACROIX).
|
|
|
|
male TYPE of Pseudolucanus groulti PLANET,
1897, from India (adapted from Araya, 2001). |
female TYPE of Pseudolucanus groulti PLANET,
1897, from India (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from
Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Lucanus
(Pseudolucanus) imitator BOUCHER & HUANG, 1991.
Lucanus
imitator BOUCHER & HUANG, 1991:32, 36 [China : Xizang = Thibet : Tangmai].
Lucanus
imitator BOUCHER & HUANG; Maes, 1992:21.
Pseudolucanus
bicolor SCHENK, 1996:508-512.
Pseudolucanus
tricolor SCHENK, 1996:508-512 (lapsus).
·
China : Xizang = Thibet : Tangmai, 30°02N-95°01E (TYPUS of Lucanus
imitator BOUCHER & HUANG, 1991).
·
China : Xizang = Thibet : Zhamo
Lucanus
(Pseudolucanus) kerleyi BOUCHER, 1994.
Lucanus
groulti PLANET; Arrow, 1950:55.
Lucanus
kerleyi BOUCHER, 1994:512-513 [India : Bhawali; Nepal : Silgarhi-Doti].
Lucanus
kerleyi BOUCHER; Araya, 1996:16.
·
India : Uttar Pradesh : Bhawali = Bhowali = Bawali, 29°22N-79°31E :
Kunayim (TYPUS of Lucanus kerleyi BOUCHER, 1994).
·
Nepal : Silgarhi-Doti : Dhalaum Area, 29°14N-80°58E (TYPUS of Lucanus
kerleyi BOUCHER, 1994).
Lucanus (Pseudolucanus) lesnei (PLANET, 1905).
Pseudolucanus lesnei PLANET, 1905:212 [China : Yunnan : Se-Tchouen :
Thibet].
Pseudolucanus lesnei PLANET; Arrow, 1950:60, pl.4
Pseudolucanus lesnei PLANET; Didier & Séguy, 1953:27
Lucanus
lesnei PLANET; Benesh, 1960:143
Lucanus lesnei PLANET; Kurosawa, 1970
Lucanus lesnei PLANET; De Lisle, 1975:203
Lucanus lesnei PLANET; Kurosawa, 1978:143
Lucanus lesnei PLANET; Kikuta, 1986:134-135
Lucanus lesnei PLANET; Boucher & Huang, 1991:31
Lucanus lesnei PLANET; Maes, 1992:21
Lucanus lesnei PLANET; Mizunuma & Nagai, 1994:211.
Lucanus lesnei (PLANET); Araya, 2001:17, figs.57-58.
·
Bhutan : Decchi Paka
·
Myanmar = Burma :
Mishmi Hills
·
China : Yunnan :
Se-Tchouen : Xizang = Thibet, 1904, col. Lieutenant Grillières (male
TYPE of Pseudolucanus lesnei PLANET, 1905 in Museum National d’Histoire
Naturelle, Paris).
|
|
|
male TYPE of Pseudolucanus lesnei PLANET,
1905, from China : Yunnan : Se-Tchouen : Xizang = Thibet (adapted from Araya,
2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus (Pseudolucanus) mazama (LECONTE, 1861).
Dorcus mazama LECONTE, 1861:345 [N. Mexico].
Parry,
1864:51; Parry, 1870:72, pl.1; Fuchs, 1882:50, pl.; Bates, 1889:382, 392;
Planet, 1895:145; Planet, 1899:19, pl.2; Villada, 1901; Planet, 1902:pl.2
Lucanus mazama ab. bicostatus ANGELL, 1916:70 [New Mexico : Ft. Wingate] (For
Poole & Gentili, 1996:297 this is a valid species).
Villada,
1923; Van Dyke in Essig, 1926:447; Tanner, 1927:32, pl.11; Benesh, 1944;
Blackwelder, 1944:195; Didier & Séguy, 1953:pl.13; Benesh, 1960:144-145;
Blackwelder, 1982:195; Boucher & Huang, 1991:38; Maes, 1992:21; Mizunuma
& Nagai, 1994:212, pl. 7; Poole & Gentili, 1996:297.
Pseudolucanus mazama (LECONTE); Reyes-Castillo, 2002:486.
USA : Utah, Arizona [Cochise Co.
(Portal--31°50N-109°10W, Miller Canyon), Madera Canyon, Oak Creek Canyon],
Colorado, New Mexico [Ft. Wingate--35°30N-108°30W];
Mexico* : Sonora (Santa
Clara--31°45N-114°30W), Chihuahua.
Bibliography
:
REYES-CASTILLO
P. (2002)
27. Lucanidae. Pp. 485-492. En LLORENTE
BOUSQUETS J. & MORRONE J.J. Biodiversidad, Taxonomía y Biogeografía de
Artrópodos de México : Hacia una síntesis de su conocimiento. Universidad
Nacional Autónoma de México. Vol. III:692 pp.
Lucanus (Pseudolucanus) oberthuri (PLANET, 1896).
Pseudolucanus oberthuri PLANET, 1896:229 [Sikkim].
Pseudolucanus oberthuri PLANET; Planet, 1899:13, pl.1
Pseudolucanus oberthuri PLANET; Planet, 1901:250
Pseudolucanus oberthuri PLANET; Planet, 1902:134, pl.1
Pseudolucanus oberthuri PLANET; Arrow, 1950:59
Pseudolucanus oberthuri PLANET; Didier & Séguy, 1952:pl.11
Pseudolucanus oberthuri PLANET; Didier & Séguy, 1953:64, 84
Pseudolucanus oberthuri PLANET; Benesh, 1960:145
Pseudolucanus oberthuri PLANET; Kurosawa, 1970
Pseudolucanus oberthuri PLANET; De Lisle, 1975
Pseudolucanus oberthuri PLANET; Kurosawa, 1978:143
Pseudolucanus oberthuri PLANET; Kikuta, 1986:134
Pseudolucanus oberthuri PLANET; Maes, 1992:21
Pseudolucanus oberthuri PLANET; Bartolozzi & Sforzi, 1994:113, 114
Pseudolucanus
oberthuri PLANET; Boucher, 1994:510-512
Lucanus oberthueri (PLANET); Mizunuma & Nagai, 1994:211, pl. 7
Pseudolucanus
oberthuri PLANET; Schenk, 1996:510.
Lucanus oberthueri (PLANET); Araya, 2001:18-19, fig.65.
·
India : Sikkim (male TYPUS of Pseudolucanus oberthuri PLANET, 1896
in Museum National d’Histoire Naturelle, Paris).
·
India : Sikkim : Lingtu
·
India : Sikkim :
Paramtsin
·
India : Sikkim : Mt. Kanchenjunga, VI-1990 (2 males, 1 female citated by
Mizunuma & Nagai 1994).
·
Nepal : Kaski Distr. : Gandrung - Ghorapani : Banthanti
|
|
|
male TYPUS of Pseudolucanus oberthuri PLANET,
1896, from Sikkim (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from
Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Lucanus
(Pseudolucanus) prometheus BOUCHER & HUANG, 1991.
Boucher
& Huang, 1991:32; Maes, 1992:21.
China :
Xizang = Thibet : Bomi-Tede*.
Lucanus
(Pseudolucanus) wittmeri (LACROIX, 1983) [Pseudolucanus].
Pseudolucanus
wittmeri LACROIX, 1983:7 [Pakistan:Sivat : Miandam].
Pseudolucanus
wittmeri LACROIX; Lacroix, 1988:7
Pseudolucanus
wittmeri LACROIX; Bartolozzi & Sforzi, 1994:114-115
Lucanus
wittmeri (LACROIX); Mizunuma & Nagai, 1994:214, pl.11, 138
Pseudolucanus
wittmeri LACROIX; Schenk, 1996:510.
Lucanus
wittmeri (LACROIX); Araya, 2001:22, figs.87-89.
·
Pakistan : Sivat : Miandam (male HOLOTYPUS of Pseudolucanus wittmeri
LACROIX, 1983).
·
Pakistan : Kalam,
2000-2400 m, 6-VI-1978, ex. coll. S. Boucher (male PARATYPUS of Pseudolucanus
wittmeri LACROIX, 1983 in Museum National d’Histoire Naturelle, Paris).
Nota : esta especie no se encuentra en
Nepal.
|
|
|
male PARATYPUS of Pseudolucanus wittmeri
LACROIX, 1983, from Pakistan : Kalam (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.