Gnaphaloryx
(Gnaphaloryx) BURMEISTER, 1847
(G. = Gnaphaloryx opacus)
= Odontaegus KRIESCHE 1935 (G. = Aegus
dain)
Gnaphaloryx
(Gnaphaloryx) curtus KIRSCH, 1877.
Kirsch,
1877:101; Albers, 1885:233; Benesh, 1960:107; Franciscolo, 1984:510; Maes,
1992:96.
Lesser
Sunda Is. : Aru I.--5°59S-134°30E;
New
Guinea : Dorei;
Bismarck
Is. : New Britain I.--5°44S-150°41E (Valoka), Duke of York I. (Manuan).
Note :
could be a ssp. of opacus.
Gnaphaloryx
(Gnaphaloryx) dain KRIESCHE, 1920 [Aegus, Odontaegus].
Kriesche,
1920:101; Kriesche, 1935:174; Didier & Séguy, 1953:156; Benesh, 1960:107;
Franciscolo, 1984:510; Maes, 1992:96.
New
Guinea : Bismarck Is. : New Britain I.--5°44S-150°41E = Neupommern I.*.
Gnaphaloryx
(Gnaphaloryx) multicavus (MIZUNUMA, 1994).
Dorcus
multicavus MIZUNUMA in MIZUNUMA & NAGAI, 1994:30-31, fig. 86(a-f), p. 274, pl.
113, fig. 427(1-5) [Pulu Pulu] [multicavatus].
Sulawesi
: Palopo, 2°55S-120°06E : Rantepao : Pulu Pulu, 30-VIII / 2-IX-1986, leg. N.
Nishikawa (3 males PARATYPUS, 2 females PARATYPUS in col. H. Fujita, Japan).
Sulawesi
: Palopo, 2°55S-120°06E : Rantepao : Pulu Pulu, I-1989 (female ALLOTYPUS in
col. T. Mizunuma, Japn).
Sulawesi : Palopo, 2°55S-120°06E : Rantepao
: Pulu Pulu, II-1989 (male HOLOTYPUS in Kanagawa Prefectural Museum, Japan) (18
males PARATYPUS, 15 females PARATYPUS in col. T. Mizunuma, Japan) (1 male
PARATYPUS, 1 female PARATYPUS in col. T.
Mizukami, Japan).
Sulawesi : Palopo, 2°55S-120°06E : Puncak,
29-III-1981, leg. H. Detani (1 male PARATYPUS in col. T. Mizunuma, Japan).
Bibliography.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Gnaphaloryx
(Gnaphaloryx) opacus BURMEISTER, 1847.
Dorcus bonasus DEJEAN 1837:194 [nomen nudum] [Java].
Lucanus
(Macrognathus) taurus FABRICIUS; Hope & Westwood, 1845 [Java].
Gnaphaloryx
opacus BURMEISTER, 1847:397 [Java].
Reiche,
1853:81; Parry, 1864:90; Vollenhoven, 1865:154, pl.2
Gnaphaloryx
rugosus ALBERS, 1885:236.
Gnaphaloryx
brevis (KIRSH) Albers, 1885:236 (nomen nudum).
Albers,
1889:236; Heyne & Taschenberg, 1908:55, pl.8; Oberthur & Houlbert,
1914:158
Gnaphaloryx
andamanus KRIESCHE, 1920:101 [Andaman].
Gnaphaloryx
tonkinensis KRIESCHE, 1920:101 [Annam : Phuc-Son; Tonkin : Than-Moi].
Nagel,
1925:3
Gnaphaloryx
burmeisteri NAGEL, 1926:119 (new name for taurus).
Nagel,
1927:153; Miwa, 1933:365, pl.3; Miwa, 1934:323; Arrow, 1935:113; Arrow, 1950:76,
pl.15; Didier & Séguy, 1953:151; Benesh, 1960:107-108; De Lisle, 1972;
Klausnitzer, 1982:33; Franciscolo, 1984:507, 509; Bomans, 1991:145; Maes,
1992:96; Kawano, 1993:13; Kawano, 1994:9
Gnaphaloryx
opacus BURMEISTER; Mizunuma & Nagai, 1994:296, 297-298, pl.134-135,
fig.631(1-48).
Gnaphaloryx
opacus BURMEISTER; Araya, 1994:6.
Gnaphaloryx
opacus BURMEISTER; Bomans & Benoit, 2005:292.
Ø Myanmar = Burma;
Ø Andaman Is.;
Ø Nicobar Is.;
Ø Taiwan : ?
(Kotosho);
Ø Thailand : Tak
(road Mae-Sot--Umphang);
Ø Vietnam : Tonkin
(Than-Moi), Annam (Phuc-Son);
Ø Malaya : Pahang
(Cameron Highlands);
Ø Borneo : Sarawak
(Songei Tutan--Kalabit Country--Mt. Poi, Kuching), Sabah (Crocker Range near
Keningau), W. Kalimantan (Ulac--Mt. Kenepai, Mt. Bawang near Pontianak);
Ø Sumatra : Simeulue I. = Simalur I., N. Sumatra Prov.
(Medan), Nias I., S. Sumatra Prov. (Mt. Dempo), Lampung Prov.;
Ø Java : W. Java
(Mt. Gede), E. Java (Mt. Argopuro), Buitenzorg, Mt. Poentjak, Mt. Salak,
Buitenzorg, Mt. Adjoeno, Toegoe, Préanger, Pengalengan, Kangean I.--
6°50S-115°20E;
Ø Java (TYPUS
perhaps in Halle Museum, Germany).
Ø
Philippines : Luzon I. : Mt. Pulog, III-1985 (1 male, 1 female in coll. Mizunuma, Japan).
Ø Philippines : Marinduque
I., 13°24N-121°57E, I-1985 (1 male in coll. Mizunuma, Japan).
Ø Philippines : Negros
I. : Mt. Canlaon, 10°22N-123°07E, VI-1990 (1 male, 1 female in coll. Mizunuma,
Japan).
Ø
Philippines : Palawan
I. : Puerto Princesa, 9°39N-118°44E
Ø Philippines : Palawan
I. : Brooke's Point, 8°54N-118°00E
Ø
Sulawesi = Celebes :
Tondano--1°14N-124°51E, Sangihe I. (Tahuna);
Ø
Moluccas Is. : Ternate
I.--0°50N-127°15E, Bachan I., Mandioli I.--0°44S-127°15E, Kasiruta
I.--0°27S-127°13E, Tidore I.--0°43N-127°25E (Bukit Gambir), Seram
I.--3°05S-129°20E = Ceram I. (Wahai), Waigeo I.--0°07S-130°50E;
Ø Lesser Sunda Is. :
Tanimbar I.--7°31S-131°30E = Yamdena I. (Saumlaki);
Ø New Guinea : Papua
(Bulolo, Fly River, Kokoda--9°04S-147°49E), Ramoi, Dorei-Hum, Andai, Korido,
Bismarck Is. : New Britain I.--5°44S-150°41E (Gazel Peninsula).
Note :
Perhaps the species could be divide into several ssp. : ssp. tonkinensis
: from Thailand to Vietnam and Taiwan; ssp. andamanus from Andaman Is.
and Nicobar Is.; ssp. ? from Malaya and Sumatra; ssp. opacus = bonasus =
taurus (mandibles with the tooth close of terminal fork) from Borneo and
Java; ssp. ? from Philippines; and, ssp. rugosus = brevis from Sulawesi
to New Guinea.
Bibliography.
BOMANS
H.E. & BENOIT P. (2005) Inventaire provisoire commenté des Coléoptères
Lucanides de l’Archipel des Philippines (98eme Contribution a l’étude des
Coléoptères Lucanides). Lambillionea, CV(2):283-301.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Gnaphaloryx (Gnaphaloryx) pollinosus DE LISLE, 1973.
De Lisle, 1973:141; Maes, 1992:96; Mizunuma
& Nagai, 1994:309, pl.147; Bomans & Bartolozzi, 1996:220.
Solomon
Is. : Bougainville I.--6°05S-155°14E (Kieta*).
Gnaphaloryx (Gnaphaloryx) stigmatifer DE LISLE, 1974.
De Lisle, 1974:786; Maes, 1992:96; Bomans &
Bartolozzi, 1996:220.
Solomon
Is. : Bougainville I.--6°05S-155°14E (Mt. Balbi*).