Lamprima LATREILLE, 1806 (G

Aegus (Gnaphaegus) MAES, 1992

(G. = Aegus squalidus)

Aegus (Gnaphaegus) albertisi BOMANS, 1992.

Bomans, 1992:181-182, 195.

New Guinea : Papua (Fly River*).

Aegus (Gnaphaegus) borneensis (NAGEL, 1926) [Gnaphaloryx].

Nagel, 1926:301; Nagel, 1927:152; Arrow, 1935:113; Benesh, 1960:100; Maes, 1992:97.

Borneo : Sarawak (Mt. Dulit*).

Aegus (Gnaphaegus) capreolus (BOILEAU, 1903).

Gnaphaloryx capreolus BOILEAU, 1903:109 (Borneo: Pontianak).

Benesh, 1960:107; Franciscolo, 1984:512; Maes, 1992:96

Gnaphaloryx capreolus BOILEAU; Mizunuma & Nagai, 1994:297, 309, pl.133-134, fig.627(1-13), pl.146, fig.93.

Gnaphaloryx capreolus BOILEAU; Bomans & Benoit, 2005:292.

Ø ? Malaya

Ø Borneo : Sabah : Mt. Trus Madi, IV-1990 (1 male in coll. Mizunuma, Japan).

Ø Borneo : Sabah : Crocker Range : near Keningau, 1000-1400 m, 6/10-III-1988 (1 male in coll. Mizunuma, Japan).

Ø Borneo : Sabah : Crocker Range : near Keningau, 1000-1400 m, 1/5-V-1988 (2 males in coll. Mizunuma, Japan).

Ø Borneo : Sabah : Crocker Range : near Keningau, 1000-1400 m, 26/29-V-1988 (1 female in coll. Mizunuma, Japan).

Ø Borneo : W. Kalimantan : Pontianak (male TYPUS in MNHN, Paris).

Ø Borneo : W. Kalimantan : Mt. Bawang, IX-1990 (1 male in coll. Mizunuma, Japan).

Ø Borneo : W. Kalimantan : Mt. Bawang, X-1990 (1 male, 1 female in coll. Mizunuma, Japan).

Ø Sumatra : W. Sumatra Prov. : Padang, II-1991 (1 male in coll. Mizunuma, Japan).

Ø Sumatra : ? : Sibolangit

Ø Philippines : Palawan I. : Brooke's Point, 8°54N-118°00E, XII-1987 (1 female in coll. Mizunuma, Japan).

Ø Philippines : Palawan I. : Brooke's Point, 8°54N-118°00E, 1987 (1 male in coll. Mizunuma, Japan).

Ø Philippines : Palawan I. : Brooke's Point, 8°54N-118°00E, I-1991 (1 male in coll. Mizunuma, Japan).

Ø Philippines : Palawan I. : Brooke's Point, 8°54N-118°00E, VII-1993 (1 male in coll. Mizunuma, Japan).

Bibliography :

BOMANS H.E. & BENOIT P. (2005) Inventaire provisoire commenté des Coléoptères Lucanides de l’Archipel des Philippines (98eme Contribution a l’étude des Coléoptères Lucanides). Lambillionea, CV(2):283-301.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Aegus (Gnaphaegus) dilaticollis (PARRY, 1864) [Gnaphaloryx] [Archip. Ind.?].

Parry, 1864:51; Benesh, 1960:101; Bomans, 1967:315; Maes, 1992:97; Mizunuma & Nagai, 1994:296, 309, pl.133.

?India;

Malaya : Pahang (Cameron Highlands);

Borneo : Sabah (Crocker Range 16 mi NW Keningau);

Sumatra : S. Sumatra (Mt. Dempo).

Aegus (Gnaphaegus) furfuraceus DE LISLE, 1970.

De Lisle, 1970:113; Maes, 1992:97; Mizunuma & Nagai, 1994:309, pl.147.

New Guinea : Mt. Wandammen--Wasion*.

Aegus (Gnaphaegus) haddeni (BENESH, 1950).

Gnaphaloryx haddeni BENESH, 1950: 49-51 (Kabasalan).

Benesh, 1960:107; Franciscolo, 1984:510; Bomans, 1990:171-172; Maes, 1992:97

Gnaphaloryx haddeni BENESH; Mizunuma & Nagai, 1994:296, 309, pl.133, fig.624(1-2).

Gnaphaloryx haddeni BENESH; Bomans & Benoit, 2005:292.

Ø Philippines : Luzon I. : Mt. Pulog, V-1985 (1 male in coll. Mizunuma, Japan).

Ø Philippines : Negros I., 10°04N-123°02E, VIII-1991 (1 male in coll. Mizunuma, Japan).

Ø Philippines : Mindanao I. : Zamboanga, 6°55N-122°04E : Kabasalan, col. F.C. Hadden (male HOLOTYPUS in California Academy of Sciences, San Francisco, USA).

Bibliography :

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Aegus (Gnaphaegus) horridus ssp. horridus BENESH, 1950.

Aegus horridus BENESH, 1950:17-18 (Nueva Vizcaya).

Benesh, 1950:49, 51; Benesh, 1960:107; Franciscolo, 1984:510; Maes, 1992:97;

Gnaphaloryx horridus ssp. horridus (BENESH); Mizunuma & Nagai, 1994:43, 296-297, pl. 133, fig.625(1-4).

Gnaphaloryx horridus (BENESH); Bomans & Benoit, 2005:292.

Ø Philippines : Luzon I. : Nueva Vizcaya, B. Benesh Collection No. 5240 (Male TYPUS in Field Museum of Natural History, Chicago, USA).

Ø Philippines : Luzon I. : Mountain Province, 8-VI-1987 (1 male in coll. Mizunuma, Japan).

Ø Philippines : Luzon I. : Mt. Pulog, III-1985 (1 male in coll. Mizunuma, Japan).

Ø Philippines : Luzon I., XI-1986 (1 male, 1 female in coll. Mizunuma, Japan).

Bibliography :

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Aegus (Gnaphaegus) horridus ssp. gunjii (NAGAI, 1994).

Gnaphaloryx horridus ssp. gunjii NAGAI in MIZUNUMA & NAGAI, 1994: 43, 297, pl. 133, fig. 625(5-7) [Mt. Apo].

Gnaphaloryx horridus (BENESH); Bomans & Benoit, 2005:292.

Ø Philippines : Mindanao I. : Mt. Apo, 7°08N-125°09E : North slope : Baracatan : Eagle Centre, 1100 m, 6-VIII-1985, leg. Masahiro Sakai (male HOLOTYPUS in National Science Museum, Tokyo).

Ø Philippines : Mindanao I. : Mt. Apo, 7°08N-125°09E, 1000 m, III-1977, leg. Yoshiaki Gunji (1 male PARATYPUS in col. T. Mizunuma, Japan).

Ø Philippines : Mindanao I. : Mt. Apo, 7°08N-125°09E, IV-1983 (1 male PARATYPUS in col. T. Mizunuma, Japan).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Aegus (Gnaphaegus) krieschei (NAGEL, 1928) [Gnaphaloryx, Xenostomus].

Nagel, 1928:4; Séguy, 1954:88; Benesh, 1960:102; Maes, 1992:97.

Borneo : Sarawak (Mt. Murud*).

Aegus (Gnaphaegus) miles ssp. sulawesiensis (NAGAI, 1994).

Gnaphaloryx miles ssp. sulawesiensis NAGAI in MIZUNUMA & NAGAI, 1994:43, pl. 133, fig. 626(6-9) [Palolo].

Sulawesi : Palu, 0°46S-119°49E : Palolo, II-1988 (1 male PARATYPUS in col. S. Nagai, Japan).

Sulawesi : Palu, 0°46S-119°49E : Palolo, X-1988 (2 males PARATYPUS, 2 females PARATYPUS in col. S. Nagai, Japan).

Sulawesi : Palu, 0°46S-119°49E : Palolo, XI-1988 (1 male PARATYPUS, 1 female PARATYPUS in col. S. Nagai, Japan).

Sulawesi : Palu, 0°46S-119°49E : Palolo, XII-1988 (female ALLOTYPUS in col. S. Nagai, Japan).

Sulawesi : Palu, 0°46S-119°49E : Palolo, V-1989 (3 males PARATYPUS, 2 females PARATYPUS in col. S. Nagai, Japan).

Sulawesi : Palu, 0°46S-119°49E : Palolo, II-1990 (3 males PARATYPUS, 2 females PARATYPUS in col. L. Bartolozzi, Italy).

Sulawesi : Palu, 0°46S-119°49E : Palolo, IX-1990 (22 male PARATYPUS in col. S. Nagai, Japan).

Sulawesi : Palu, 0°46S-119°49E : Palolo, 600 m, XII-1990 (male HOLOTYPUS in col. S. Nagai, Japan).

Aegus (Gnaphaegus) miles ssp. miles (VOLLENHOVEN, 1865).

Gnaphaloryx miles VOLLENHOVEN, 1865:155, pl. 2 [Halmahera].

Parry, 1870:63; Boileau, 1903:110; Arrow, 1935:113; Benesh, 1960:107; Franciscolo, 1984:506; Maes, 1992:97; Mizunuma & Nagai, 1994:43, 297, 309, pl. 133, 146.

Moluccas Is. : Halmahera I.*--0°40N-128°E, Ternate I.--0°50N-127°15E, Bachan I., Batandua I. = Mayu I., Gebe I.--0°05S-129°25E = Gebeh I.*.

Note : Mizunuma & Nagai (1994) given Ternate I. as the type locality of miles.

Aegus (Gnaphaegus) miles ssp. laticornis (BOILEAU, 1903).

Gnaphaloryx miles ssp. laticornis BOILEAU, 1903.

New Guinea* : Ramoi, Papua (Fly River*).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Aegus (Gnaphaegus) notarii ssp. gracilis (NAGAI, 1994).

Gnaphaloryx notarii ssp. gracilis NAGAI in MIZUNUMA & NAGAI, 1994:43, pl. 134, fig. 629(3-8) [Flores I.].

Lesser Sunda Is. : Sumbawa I., 8°44S-118°00E, VI/VII-1990 (3 males PARATYPUS, 1 female PARATYPUS in col. S. Nagai, Japan).

Lesser Sunda Is. : Flores I., 8°37S-121°19E : Ruteng, I-1990 (female ALLOTYPUS in col. S. Nagai, Japan).

Lesser Sunda Is. : Flores I., 8°37S-121°19E, V-1990 (male HOLOTYPUS, 8 males PARATYPUS, 3 females PARATYPUS in col. S. Nagai, Japan).

Note : The name gracilis NAGAI is junior homonym of gracilis DEYROLLE 1865.

Aegus (Gnaphaegus) notarii ssp. notarii (LACROIX, 1986).

Gnaphaloryx notarii ssp. notarii LACROIX, 1986.

Mizunuma & Nagai, 1994:43, 297, 309, pl. 134.

Note : perhaps this species is a ssp. of A. squalidus ?

Lesser Sunda Is. : Timor I.--9°13S-124°56E (Oesusu);

Sulawesi = Celebes : Todjamboe*.

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Aegus (Gnaphaegus) perforatus ssp. perforatus (RITSEMA, 1885).

Gnaphaloryx perforatus RITSEMA, 1885:39, pl. 3 [Sumatra].

Arrow, 1935:113; Benesh, 1950:18; Benesh, 1960:108; Franciscolo, 1984:507; Bomans, 1990:172; Maes, 1992:97; Mizunuma & Nagai, 1994:43-44, 296, pl. 133.

Malaya : Kedah

Sumatra*

?Borneo

Aegus (Gnaphaegus) perforatus ssp. sabanus (NAGAI, 1994).

Gnaphaloryx perforatus ssp. sabanus NAGAI in MIZUNUMA & NAGAI, 1994:43-44, pl. 133, fig. 622(4-8) [Crocker Range].

Borneo : Sarawak : Bintulu, 22-VI-1976, leg. H. Egawa (1 male PARATYPUS in col. K. Sakai, Japan).

Borneo : Sabah : 16 mi NW Keningau : Crocker Range, 1000-1400 m, III/IV-1988 (6 males PARATYPUS, 8 females PARATYPE in col. S. Nagai, Japan).

Borneo : Sabah : 16 mi NW Keningau : Crocker Range, 1000-1400 m, 11/15-IV-1988 (male HOLOTYPUS, female ALLOTYPE in col. S. Nagai, Japan).

Borneo : Sabah : 10 mi NW Keningau : Crocker Range, 1000 m, 10-V-1989, leg. S. Nagai (1 male PARATYPUS in col. S. Nagai, Japan).

Borneo : Sabah : Mt. Trus Madi, IV-1990 (1 male PARATYPUS in col. S. Nagai, Japan).

Borneo : Sabah : Mt. Terian, 1000 Ft., V-1990 (9 males PARATYPUS, 4 females PARATYPE in col. S. Nagai, Japan).

Borneo : W. Kalimantan : Pontianak.

Note Aegus (Gnaphaegus) sabanus NAGAI 1994 is junior homonym of Aegus (Aegus) sabanus NAGAI 1994.

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Aegus (Gnaphaegus) pulverosus BENESH, 1952.

Benesh, 1952:136-137; Benesh, 1960:105; Maes, 1992:97; Bomans & Bartolozzi, 1996:219.

Solomon Is. : Bougainville I.--6°05S-155°14E (Piva River*)

Aegus (Gnaphaegus) sculptipennis (PARRY, 1864) [Gnaphaloryx].

= Gnaphaloryx aper GESTRO 1881 [Fly River].

Parry, 1864:52; Parry, 1874:362, pl.3; Gestro, 1881:324; Ritsema, 1881:32; Albers, 1885:232; Arrow, 1935:113; Benesh, 1960:106; Klausnitzer, 1982:45; Franciscolo, 1984:507; Maes, 1992:97; Mizunuma & Nagai, 1994:297, 309, pl.134; Bartolozzi, 1996:129.

New Guinea* : Irian Jaya [Jayapura--2°28S-140°38E--Cyclops Mts.--Sentani], Papua (Itikinumu, Fly River--Kiunga).

Aegus (Gnaphaegus) squalidus (HOPE & WESTWOOD, 1845).

Sturm, 1826:165; Dejean, 1837:194; Hope & Westwood, 1845:19; Burmeister, 1847:397; Heyne & Taschenberg, 1908:55; Boileau, 1911:443; Oberthur & Houlbert, 1914:160; Arrow, 1935:113; Benesh, 1950:50; Benesh, 1960:107, 108; Franciscolo, 1984:507, 510; Bomans, 1990:172; Maes, 1992:97; Mizunuma & Nagai, 1994:297, 309, pl.134, 147.

Aegus (Gnaphaegus) squalidus ssp. ericsoni (BOILEAU, 1911) [Gnaphaloryx] [erichsoni].

Malaya : Perak (Taiping--4°51N-100°41E), Pahang (Cameron Highlands);

Borneo;

Sumatra : N. Sumatra Prov. (Berastagi), S. Sumatra Prov. (Sétinjak*, Tandjong Djati--Ranau--Palembang, Mt. Dempo), Lampung Prov., ? (Deli).

Aegus (Gnaphaegus) squalidus ssp. squalidus (HOPE & WESTWOOD, 1845) [Lucanus, Gnaphaloryx].

= Lucanus setosus STURM 1826 (nomen nudum).

= Dorcus lutulentus (DE HAAN) DEJEAN 1837 [nomen nudum] [Java].

= Dorcus tomentosus DEJEAN 1837 [nomen nudum] [Java].

= Lucanus (Macrognathus) vestitus (DE HAAN) HOPE & WESTWOOD 1845 (nomen nudum) [Java].

Java* : E. Java (Mt. Argopulo), Mt. Poentjak, Mt. Kawie, Mt. Ardjoeno, Toegoe, Kangean I.--6°50S-115°20E.

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