Lamprima LATREILLE, 1806 (G

Figulus W.S. MACLEAY, 1819

(G. = Lucanus striatus)

= Eudora CASTELNAU 1840 (G. = Lucanus striatus)

Figulus acutangulus ARROW, 1935.

Arrow, 1935:120; Benesh, 1953:31; Benesh, 1960:24; Bomans, 1989:13; Maes, 1992:35.

Vietnam : Tonkin (Hoa-binh*).

Figulus albertisi GESTRO, 1881.

Gestro, 1881:335; Benesh, 1960:24; Maes, 1992:35.

New Guinea : Papua (Fly River*), Ramoi*.

Figulus andamanus KRIESCHE, 1920.

Kriesche, 1920:106; Arrow, 1950:223; Benesh, 1960:24; Maes, 1992:35.

Andaman Is.* : Chatham.

Figulus angustus BOMANS, 1989.

Bomans, 1989:15-16.

India : West Bengal (Calcutta*).

Figulus approximatus BENESH, 1955.

Benesh, 1955:61; Benesh, 1960:154; Vinson, 1967:321; Maes, 1992:35.

Mauritius I.--20°17S-57°34E = Isle of France*.

Figulus aratus ARROW, 1935.

= Figulus parvulus (BOILEAU) DIDIER 1937.

Arrow, 1935:119; Arrow, 1938:61, pl.4; Arrow, 1950:224, pl.22; Didier, 1937; Benesh, 1960:24; Bomans, 1986:8; Bartolozzi & Bomans, 1988:80; Maes, 1992:36.

India : West Bengal (Calcutta), Tamil Nadu (Coimbatore, Nilgiri Hills*Hattikeri*, Anamalai Hills, Coorg);

Sri Lanka = Ceylon : North Central Prov. [Polonnaruwa Distr. (Alut Oya)].

Saproxylophagous : Arecaceae.

Figulus beccarii GESTRO, 1881.

Gestro, 1881:338; Benesh, 1960:24; Weinreich, 1971; Weinreich, 1972; Maes, 1992:36.

Sumatra : N. Sumatra Prov. (Berastagi, Mt. Gunung), W. Sumatra Prov. (Mt. Singalan*, Fort de Kock).

Figulus bicolor BOMANS, 1986.

Bomans, 1986:10, pl.1; Maes, 1992:36.

Vietnam : Tonkin (Thatkhe*, Bao Lac*).

Figulus binodulus WATERHOUSE, 1873.

Figulus binodulus WATERHOUSE, 1873:227.

Lewis, 1883:339; Nagel, 1928:296; Miwa, 1934:320, pl.4; Benesh, 1960:24; Nomura, 1960:43; Bomans, 1986:11; Fujita & Ichikawa, 1986:26; Isida & Fujioka, 1988:2; Bomans, 1989:16;

Figulus binodorius WATERHOUSE; Araya, 1991

Abe, Kudoh, Ichikawa & Saitoh, 1992:218; Araya, Obuchi & Ochi, 1992:31, 34; Maes, 1992:36; Araya & Obuchi, 1993:43-47; Sakaino, Yu & Chu, 1994:122; Mizunuma & Nagai, 1994:299, pl. 136; Araya & Obuchi, 1994:28, 30;

Figulus binodulus WATERHOUSE; Araya & Takehiro, 1994

Araya, 1994:3; Araya, 1996:21

Figulus binodulus WATERHOUSE; Cammaerts & Bomans, 1997:13.

Figulus binodulus WATERHOUSE; Araya, 2001:272.

· China

· Korea

· Japan : Honshu I. : Fukui Pref. (citated by Araya, 2001).

· Japan : Honshu I. : Izu Is. : Mikurajima I.

· Japan : Honshu I. : Izu Is. : Hachijojima I., 33°07N-139°48E : Sueyoshi

· Japan : Shikoku I.

· Japan : Kyushu I.

· Japan : Konose

· Japan : Nara-shi

· Taiwan : Kosempo

· Vietnam : Tonkin

Bibliography.

ARAYA K. (1991) Notes on the biology of Figulus binodorius. Anim. Mag. Nat. Hist., 225:49-51.

ARAYA K. & TAKEHIRO O. (1994) Life histories of Japanese Lucanid Beetles (Coleoptera : Lucanidae). IV. Figulus MAC LEAY, 1819 (Part 2). Mushi Series, 29(5):28-32.

ARAYA K. (2001) New records of Ceruchus lignarius monticola Nakane, 1978 (Coleoptera, Lucanidae) from Fukui Prefecture, Central Japan, with reference to Lucanid fauna of Fukui Prefecture. Sukunahikona, Special Publication of the Japan Coleopterological Society, 1:271-273.

CAMMAERTS R. & BOMANS H.E. (1997) Quelques Lucanidae de Nouvelle-Guinée dont Penichrolucanus leveri Arrow, une nouvelle espèce de Figulus MacLeay et la larve de Figulus sulcicollis Hope & Westwood (Coleoptera). Bull. Annls. Soc. R. belge Ent., 133:3-20.

Figulus boileaui BOMANS, 1986.

Bomans, 1986:10, pl.1; Maes, 1992:36; Moore & Cassis, 1992:9.

Australia*.

Figulus boninensis NAKANE & KUROSAWA, 1953 [boniensis].

Nakane & Kurosawa, 1953:1; Benesh, 1960:154; Nomura, 1960:44; Fujita & Ichikawa, 1986:26; Kusui, 1987:28-29; Nakane, 1987:135; Yamaguchi, 1987:37; Isida & Fujioka, 1988:2; Abe, Kudoh, Ichikawa & Saitoh, 1992:218; Araya, Obuchi & Ochi, 1992:34; Maes, 1992:36; Mizunuma & Nagai, 1994:299, pl. 136; Araya & Obuchi, 1994:31; Araya, 1996:21.

Japan : Tokyo Met. [Bonin Is.*--28°N-142°E = Ogasawara Is. (Mukoujima I., Chichijima I., Baijima I., Otoutojima I., Anijima I., Hahajima I.)].

Figulus borneensis NAGEL, 1941.

Nagel, 1941:71; Benesh, 1960:24; Bomans, 1987:13; Maes, 1992:36; Mizunuma & Nagai, 1994:300, pl. 136.

Borneo : W. Kalimantan (Mt. Bawang near Singkawang), S. Kalimantan (Bandjermasin*).

Figulus caecus BOMANS, 1988.

Bomans, 1988:14; Maes, 1992:36; Mizunuma & Nagai, 1994:314.

New Guinea : Papua (Okapa--Ibusa*, Ialibu*, Mt. Michael--Lufa*, Kassem Pass*).

Figulus cambodiensis DEYROLLE, 1874.

Deyrolle, 1874:414; Arrow, 1950:221; Benesh, 1960:24; Bomans, 1986:9; Bomans & Lacroix, 1989:24; Maes, 1992:36.

Cambodia*.

Figulus canaliculatus NAGEL, 1941.

Nagel, 1941:69; Benesh, 1960:24; Bomans, 1988:7; Maes, 1992:36.

New Guinea* : Papua (Wau--7°19S-146°37E, Bulolo River, Madang--5°12S-145°43E, Korker I.).

[Figulus capensis (CASSTROM en THUNBERG, 1781) [Lucanus, Platycerus] Tenebrionidae of South Africa (BOMANS, 1986)].

Casstrom en Thunberg, 1781; Brinck, 1956:305; Benesh, 1960:24; Bomans, 1986:8; Maes, 1992:36.

Figulus caviceps BOILEAU, 1902.

Figulus arrowi NAGEL 1941 [Viet Nam : Tonkin : Rivière Claire, Hoa Binh].

Boileau, 1902:205; Nagel, 1941:70; Arrow, 1950:223, pl. 22; Didier & Séguy, 1953:180; Benesh, 1960:24, 25; Bomans, 1986:8; Bomans, 1987:10; Bomans, 1988:15; Bomans & Lacroix, 1989:25; Maes, 1992:36; Bartolozzi & Sforzi, 1994:108-109, 114; Mizunuma & Nagai, 1994:299, pl. 136.

India : West Bengal (Darjeeling), Madhya Pradesh (Supkhar, Balaghat);

Nepal : Chitwan Nat. Park--Sauraha, 9 m. W of Hitaura;

Bhutan: Pedong;

Myanmar = Burma : ? (Teinzo*);

Thailand : Chiang Mai (Chiang Mai--18°50N-98°57E, Mt. Doi Pui);

Vietnam: Tonkin (Rivière Claire, Hoa Binh).

Figulus cavifrons BOMANS, 1987.

Bomans, 1987:6, pl.6; Maes, 1992:36.

Sumatra : N. Sumatra Prov. (Dolok Merangir), S. Sumatra Prov. (Benkoelen--Maeara Tenam*), ? (Langsa).

Figulus cheesmani ARROW, 1938 [cheesmanae].

Arrow, 1938:61, pl.4; Benesh, 1960:25; Maes, 1992:36.

New Guinea : Papua (Kokoda*--9°04S-147°49E).

Figulus cicatricosus BOILEAU, 1905.

Boileau, 1905:28; Arrow, 1950:225, pl.22; Benesh, 1960:25; Bomans, 1986:11; Maes, 1992:36.

India : Tamil Nadu (Nilgiri Hills).

Figulus cochinchinensis NAGEL, 1928.

Nagel, 1928:291; Benesh, 1960:25; Bomans, 1986:11; Maes, 1992:36.

Vietnam : Cochinchina*.

Figulus concatenatus ARROW, 1938.

Arrow, 1938:59, pl.4; Benesh, 1960:25; Bomans, 1986:11; Maes, 1992:36.

Borneo : Sarawak (Mt. Matang*).

Figulus confusus WESTWOOD, 1838.

Figulus striatus W.S. MACLEAY (nec OLIVIER) 1819.

Figulus punctatostriatus DEYROLLE 1874 [Malacca, Timor I.].

Figulus punctatostriatus var. malaccae (ZANG) KRIESCHE 1922 [Singapore].

W.S. Macleay, 1819:100; Westwood, 1838:262; Burmeister, 1847:439; Deyrolle, 1874:413; Kriesche, 1922:133; Benesh, 1950:52-53; Benesh, 1960:25; Bomans, 1986:9; Maes, 1992:36.

Cambodia;

Malaya = Malacca : Singapore;

Lesser Sunda Is. : Timor I.--9°13S-124°56E, Larat I.;

Caroline Is. : Palau I.--7°30N-134°34E (Garakayo).

Figulus coomani ARROW, 1935.

Arrow, 1935:120; Benesh, 1960:25; Maes, 1992:37.

Vietnam* : Tonkin (Backan = Backau*, Hoa-binh*).

Figulus corvinus KRIESCHE, 1922.

Kriesche, 1922:129; Benesh, 1960:25; Bomans, 1987:10; Maes, 1992:37.

Cameroon : Moliwe near Victoria*.

Figulus crupsinus BOMANS, 1987.

Figulus crupsinus BOMANS, 1987:9, pl.6 (Luzon).

Figulus cruspinus BOMANS; Maes, 1992:37.

Figulus crupsinus BOMANS; Bomans & Benoit, 2005:291.

Philippines : Luzon I. : North Luzon, 1984, leg. Whitehead (male HOLOTYPUS in coll. H. Bomans in Natural History Museum, London).

Bibliography :

BOMANS H.E. & BENOIT P. (2005) Inventaire provisoire commenté des Coléoptères Lucanides de l’Archipel des Philippines (98eme Contribution a l’étude des Coléoptères Lucanides). Lambillionea, CV(2):283-301.

Figulus curvicornis BENESH, 1950.

Figulus curvicornis BENESH, 1950::51, 53, 56-57, pl.1 (Mt. Makiling).

Benesh, 1960:25; Bomans, 1987:10; Maes, 1992:37; Sakaino & Ochi, 1993:20-22; Sakaino, Yu & Chu, 1994:122

Figulus curvicornis BENESH; Mizunuma & Nagai, 1994:299-300, pl. 136, fig.646 (1-2).

Figulus curvicornis BENESH; Bomans & Benoit, 2005:291.

Ø Taiwan : Lanyu I., 22°00N-121°32E : Lan Hsu I. = Botel Tobago I., 10-IV-1977 (1 ex in coll. Mizunuma, Japan).

Ø Philippines : Luzon I. : Laguna : Mt. Makilling, 8/9-I-1931, coll. V. Madrid (HOLOTYPUS in California Academy of Sciences, San Francisco, USA).

Ø Philippines : Luzon I. : Camarines Sur : Mabatobato Pili*

Ø Philippines : Luzon I. : Rizal Province : Marikima*

Ø Philippines : Luzon I. : Asin Spa : Benget

Ø Philippines : Sibuyan I., 12°25N-122°35E, IX-1983 (1 ex in coll. Mizunuma, Japan).

Bibliography :

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Figulus daitoensis FUJITA & ICHIKAWA, 1986 [daitonensis].

Figulus punctatus SAKAINO (nec WATERHOUSE) 1983.

Sakaino, 1983:35; Fujita & Ichjikawa, 1986:24-31; Isida & Fujioka, 1988:2; Maes, 1992:37; Sakaino & Ochi, 1993:21; Mizunuma & Nagai, 1994:pl. 136; Araya & Obuchi, 1994:28-29; Araya, 1996:21.

Japan : Okinawa Pref. [Kitadaitojima I.*, Minamidaitojima I. (Daitô-jinja*)].

Figulus decorus DE LISLE, 1968.

De Lisle, 1968:397; Maes, 1992:37.

New Guinea : Bismarck Is. : Mussau I.--1°26S-149°36E (Talumalaus*), Mioko I. (New Larrenburg).

Figulus deletus BOMANS, 1989.

Bomans, 1989:17, 18.

China : Guangxi*;

Vietnam : Tonkin (Than Moi*, Cho Ganh*, Hoah Binh*, Tuyen Kan*), Annam (Phuc Son*).

Figulus delicatus BOMANS, 1986.

Bomans, 1986:11; Maes, 1992:37.

Malaya : Singapore*.

Figulus delislei BENESH, 1953.

Benesh, 1953:30; Benesh, 1960:25; Maes, 1992:37.

Vietnam : Tonkin (Hoa Binh*).

Figulus detzneri KRIESCHE, 1922.

Kriesche, 1922:136; Benesh, 1960:25; Maes, 1992:37.

New Guinea : Papua (Sattelberg*).

Figulus elateroides BOMANS, 1986.

Figulus sp. Bomans, 1967, 1968, 1970.

Bomans, 1967:317-323; Bomans, 1968:31-42; Bomans, 1970:232; Bomans, 1986:10, pl.1; Maes, 1992:37.

Laos : Vientiane*, Khong Sedone*.

Figulus excavatus BOMANS, 1986.

Bomans, 1986:11; Bomans, 1988:7; Maes, 1992:37.

New Guinea : Papua (Northern District--Managalase Plateau*).

Figulus exquisitus BOMANS, 1987.

Bomans, 1987:6, pl.6; Maes, 1992:37.

Malaya : Singapore*.

Figulus fissicollis FAIRMAIRE, 1849.

Figulus modestus PARRY 1862 [New Zealand, error].

Figulus monochromus DIDIER 1930 [Fidji: Viti: Rewa].

Fairmaire, 1849:414; Parry, 1862:113; Parry, 1884:60; Didier, 1930:171; Benesh, 1950:51-54, pl.1; Séguy, 1954:89; Benesh, 1958:66; Benesh, 1960:25; De Lisle, 1972; Sakaino & Ochi, 1983:1-2; Bomans, 1986:8; Bomans, 1986:7; Maes, 1992:37; Sakaino & Ochi, 1993:20; Sakaino, Yu & Chu, 1994:122; Mizunuma & Nagai, 1994:pl. 136.

Taiwan : Lanyu I.--22°00N-121°32E (Lan Yu I. = Botel Tobago I.);

Philippines : Luzon I. (Asingan--Pangasinan, Camarines Sur--Mabatobato Pili, Batangas--Mt. Lobo--Malabringo), Negros I. (Mt. Canlaon--10°22N-123°07E), Mindanao I. [Zamboanga--6°55N-122°04E (Kabasalan)];

[no Caroline Is. : Palau I., misidentification with F. lupinus];

Solomon Is. : Bougainville I.--6°05S-155°14E;

[no Marianas Is. : Guam I., confusion with F. manillarum];

Gilbert Is.--0°N-174°E;

Fidji Is. : Viti Levu I.--17°48S-177°58E (Rewa River), Vanua Levu I.--16°34S-179°13E, Taveuni I.--16°50S-179°59W;

Samoa Is.--14°S-171W;

Tonga Is. : Tongatapu I.*--21°09S-175°13W.

Note : Bomans (1986:8) considers that F. fissicollis does not exist in Philippines and there is a confusion with F. mediocris and perhaps F. lupinus.

Figulus foveicollis (BOISDUVAL, 1835).

Platycerus foveicollis BOISDUVAL, 1835:235.

Fairmaire, 1849:414

Figulus insularis BLANCHARD, 1853:142, pl.9

Figulus woodlarkianus MONTROUZIER, 1857:26 [Woodlarck I.].

Figulus lifuanus MONTROUZIER, 1857:26 [Lifu I.].

Parry, 1864:100

Figulus foveicollis subsp. gazellae KRIESCHE, 1922:132, 137 [Fidji : Viti Levu : Rawa River].

Figulus napoides KRIESCHE, 1922:132, 137 [Tahiti, Not Singapur].

Nagel, 1928:295; Arrow, 1938:52; Emden, 1952:301; Benesh, 1955:64; Benesh, 1958:65; Benesh, 1960:25; Bomans, 1989:15; Maes, 1992:37; Mizunuma & Nagai, 1994:300, pl. 136

Platycerus foveicollis BOISDUVAL; Cammaerts & Bomans, 1997:15.

New Guinea : D'Entrecasteaux Is. : Woodlarck I., 9°02S-152°43E;

Vanuatu Is., 16°54S-167°59E = New Hebrides Is.;

Loyalty Is. : Lifu I., 20°57S-167°12E;

New Caledonia I., 21°S-165°E;

Fidji Is. : Viti Levu I., 17°48S-177°58E : Rewa River

Fidji Is. : Vanua Levu I., 16°34S-179°13E

Fidji Is. : Taveuni I., 16°50S-179°59W;

Tonga Is. : Tongatapu I.*, 21°09S-175°13W;

Samoa Is., 14°S-171W;

Society Is. : Tahiti I., 17°38S-149°28W : Valley SE Tahateo Huahine

Xylophagous : Malvaceae : Hibiscus.

Bibliography.

Figulus fulgens BOMANS, 1986.

Bomans, 1986:8, pl.3; Maes, 1992:37.

Tuvalu Is. = Ellice Is.*.

Figulus fusconitens DIDIER, 1930.

Didier, 1930:172; Benesh, 1960:25; Bomans, 1986:12; Maes, 1992:37.

Sumatra : N. Sumatra Prov. : Nias I.*.

Figulus gestroi NAGEL, 1928.

Nagel, 1928:294; Benesh, 1960:26; Maes, 1992:37.

New Guinea*.

Figulus gracilentus DIDIER, 1930.

Didier, 1930:171; Benesh, 1960:26; Bomans, 1986:12; Maes, 1992:37.

Moluccas Is. : Ambon I.--3°39S-128°07E = Amboina I.*.

Figulus granulosus BOMANS, 1986 [granulosa].

Bomans, 1986:11, pl.1; Maes, 1992:37.

Sumatra : S Sumatra Prov. (Palembang*).

Figulus hoogstraali BENESH, 1955.

Figulus hoogstraali BENESH, 1955:63 (Mt. Mac Kinley).

Benesh, 1960:154; Bomans, 1987:10; Maes, 1992:37.

Figulus hoogstraali BENESH; Bomans & Benoit, 2005:291.

Ø Philippines : Mindanao I., 7°44N-125°02E : Mt. McKinley : East slope, 6400 Ft., VIII/IX-1946 (male TYPUS in coll. V. Hoogstraal in Field Museum of Natural History, Chicago, USA).

Bibliography :

Figulus hoplocnemus DE LISLE, 1974.

De Lisle, 1974:799; Maes, 1992:38.

New Guinea : Papua (Torricelli Mts.--Mokai Vill.*).

Figulus hornabrooki BOMANS, 1986.

Bomans, 1986:12; Bomans, 1987:5; Bomans, 1988:7; Maes, 1992:38.

New Guinea : Papua [Okapa--Takai*, E.H. Prov.--Kaimantu--Orenunka*, Eastern Highlands* (Okapa), E.H. Prov.--Rintobe*, Madang*--5°12S-145°43E, Morobe--7°42S-147°34E (Lae*, Kaiapit*)].

Figulus horni ZANG, 1905.

Zang, 1905:161; Arrow, 1950:222; Benesh, 1960:26; Bomans, 1986:9; Bartolozzi & Bomans, 1988:80; Maes, 1992:38.

Sri Lanka = Ceylon : North Western Prov. [Puttalam Distr. (10 mi E Puttalam)], North Central Prov. [Anuradhapura Distr. (Ritigala Nat. Reserve, Habarane)], Western Prov. [Colombo Distr. (Colombo), Gampaha Distr. (Negombo*)].

Figulus howei DE LISLE, 1967.

De Lisle, 1967:542; Bomans, 1989:14; Maes, 1992:38; Moore & Cassis, 1992:9.

Australia : Lord Howe I.*--31°33S-159°05E.

Saproxylophagous.

Figulus humeralis DIDIER, 1930.

Didier, 1930:165; Benesh, 1960:26; Maes, 1992:38.

Lesser Sunda Is. : Larat I.*;

New Guinea.

Figulus impressicollis RITSEMA, 1896.

Ritsema, 1896:139; Benesh, 1960:26; Maes, 1992:38.

Borneo : Sarawak (Kalabit Country), W. Kalimantan (Mt. Kenepai*).

Figulus incertus BOMANS, 1987.

Bomans, 1987:5, pl.5; Maes, 1992:38.

New Guinea : Papua (Redscar Bay*).

Figulus incisus BOMANS, 1986.

Bomans, 1986:7; Maes, 1992:38.

Malaya : Singapore*.

Figulus insolitus BOMANS, 1987.

Bomans, 1987:10, pl.7; Maes, 1992:38.

Bhutan : Pedong*.

Figulus insulanus BOMANS, 1986.

Bomans, 1986:7, pl.3; Maes, 1992:38.

Vanuatu Is.--16°54S-167°59E = New Hebrides Is. : Vila*.

Figulus integricollis THOMSON, 1862.

Thomson, 1862:431; Gestro, 1881:334; Didier & Séguy, 1953:58; Benesh, 1960:26; Bomans, 1987:13; Maes, 1992:38

Marianas Is.* : Guam I.--13°27N-144°46E, Rota I.--14°08N-145°14E.

Figulus interruptus WATERHOUSE, 1874 [India].

Waterhouse, 1874:7; Gravely, 1915:430; Arrow, 1950:221; Benesh, 1960:26; Bartolozzi & Bomans, 1988:81; Maes, 1992:38.

? Sri Lanka = Ceylon : Central Prov. [Kandy Distr. (Peradeniya)].

Figulus javanensis NAGEL, 1941.

Nagel, 1941:73; Benesh, 1960:26; Maes, 1992:38.

Java : G. Oengeran*.

Figulus joliveti BOMANS, 1986.

Bomans, 1986:11, pl.1; Bomans, 1984:357; Maes, 1992:38.

New Guinea : Papua (Goroka*).

Figulus kinabaluensis BOMANS, 1987.

Bomans, 1987:13, pl.8; Maes, 1992:38.

Borneo : Sabah (Mt. Kinabalu*).

Figulus laevipennis MONTROUZIER, 1860.

Montrouzier, 1860:286; Fauvel, 1903:357; Benesh, 1960:26; Bomans, 1986:9; Maes, 1992:38; Mizunuma & Nagai, 1994:298-299, pl. 136.

New Caledonia I.*--21°S-165°E (Sarramea);

Art I.

Figulus laevithorax BOMANS, 1987.

Bomans, 1987:6, pl.5; Maes, 1992:38; Moore & Cassis, 1992:9-10.

Australia : Northern Territory (Darwin--12°27S-130°50E*), Queensland (Cape York*--33°15S-150°08E).

Figulus lansbergei RITSEMA, 1880.

Ritsema, 1880:217; Benesh, 1960:26; Bomans, 1986:9; Bomans, 1987:6; Bomans, 1990:175-176; Maes, 1992:38; Mizunuma & Nagai, 1994:300, pl. 136.

Lesser Sunda Is. : Sumbawa I.*--8°44S-118°00E, Flores I.--8°37S-121°19E.

Figulus laoticus BOMANS, 1986.

Bomans, 1986:11, pl.2; Bomans, 1991:145; Maes, 1992:38; Mizunuma & Nagai, 1994:299, pl. 136.

Thailand : Chiang Mai (Chiang Mai--18°50N-98°57E, Mt. Doi Pui);

Laos : Pakse*;

Vietnam : Cochinchina*.

[Figulus laticollis (ESCHSCHOLTZ) DEJEAN 1837 [nomen nudum] [Philippines].

Dejean, 1837:194; Benesh, 1950:57; Benesh, 1960:152; Sakaino & Ochi, 1983:1-2; Maes, 1992:38.

Figulus leptochilus DE LISLE, 1974.

De Lisle, 1974:802; Maes, 1992:38; Bomans & Bartolozzi, 1996:219.

Solomon Is. : Guadalcanal I.--9°37S-160°02E (mouth of Tenaru River*).

Figulus lilliputanus WESTWOOD, 1855.

Figulus clivinoides THOMSON 1862 [Australia].

Westwood, 1855:219-220, pl.12; Thomson, 1862:432; Parry, 1863:450; Mjoberg, 1916:13; Benesh, 1955:66; Benesh, 1960:26; Klausnitzer, 1982:15; Bomans, 1986:8-9; Maes, 1992:36, 38; Moore & Cassis, 1992:10.

Australia : Queensland, New South Wales (Murray--Darling Basin--32to36°S-115to148°E), South Australia (Adelaide*--34°56S-138°36E), Western Australia (Fremantle--32°03S-115°44E).

Saproxylophagous.

Note : Bomans (1986:8) considers that F. clivinoides could be a good species from Queensland and New South Wales; F. lilliputanus is from Western Australia and South Australia.

Figulus linearis DIDIER, 1929 [Cardanus].

Didier, 1929:81; Arrow, 1943:139; Arrow, 1950:225; Benesh, 1960:26; Bomans, 1986:9; Maes, 1992:38.

India : Kerala = Malabar (Mahé*).

Figulus lozoki KRIESCHE, 1922.

Kriesche, 1922:135; Benesh, 1960:26; Maes, 1992:38.

New Guinea : Papua (Sattelberg*).

Figulus lupinus KRIESCHE, 1922.

Figulus benningseni (ZANG) BOMANS 1986.

Kriesche, 1922:131; Benesh, 1950:54; Benesh, 1955:67; Benesh, 1960:25; Bomans, 1986:8; Bomans, 1986:7-8; Maes, 1992:39.

Caroline Is. : Palau I.*--7°30N-134°34E.

Figulus magnus BENESH, 1955 [mangus].

Benesh, 1955:62; Benesh, 1960:154; Maes, 1992:39.

Seychelles Is.*--4°30S-55°30E.

Figulus manillarum HOPE & WESTWOOD, 1845.

Figulus angustatus (EISCHSCHOLTZ) Dejean, 1837:194 [nomen nudum] [Philippines].

Figulus manillarum HOPE & WESTWOOD, 1845:26 [Philippines].

Fairmaire, 1849:415; Parry, 1864:100

Figulus lilliputanus (WESTWOOD); Zimmerman, 1942.

Benesh, 1950:51-54, 57, pl.1; Benesh, 1960:26; De Lisle, 1972; Bomans, 1986:8; Maes, 1992:39; Fujita, 1994:15

Figulus manillarum HOPE & WESTWOOD; Mizunuma & Nagai, 1994:300, pl. 136, fig.656(1).

Figulus manillarum HOPE & WESTWOOD; Bomans & Benoit, 2005:291.

Ø Philippines : Luzon I. : Manilla, 14°35N-120°57E, col. Cuming (TYPUS in Natural History Museum, London).

Ø Philippines : Luzon I. : Alabang

Ø Philippines : Luzon I. : Camarines Sur : Mabatobato Pili

Ø Philippines : Luzon I. : Laguna : Mt. Makilling

Ø Philippines : Mindoro I., 12°58N-121°03E : San José

Ø Philippines : Sibuyan I., 12°25N-122°35E

Ø Philippines : Leyte I. : Tacloban, 11°15N-124°59E

Ø Philippines : Mindanao I. : Zamboanga, 6°55N-122°04E : Kabasalan

Ø Philippines : Palawan I., 9°37N-118°32E, 21-XII-1978 (1 ex. In coll. Mizunuma, Japan).

Ø Marianas Is. : Guam I., 13°27N-144°46E.

Bibliography :

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

Figulus marginalis RITSEMA, 1879.

Figulus subcastaneus VOLLENHOVEN 1865 (pre-ocupado por subcastaneus Westwood).

Vollenhoven, 1865:146, 156; Ritsema, 1879:189; Ritsema, 1895:138; Sharp & Muir, 1912:575; Oberthur & Houlbert, 1914:223; Benesh, 1960:26; Bomans, 1986:9; Bomans, 1988:14; Bomans, 1989:13; Bomans, 1990:176; Maes, 1992:39; Mizunuma & Nagai, 1994:314.

Borneo* : W. Kalimantan (Pontianak), ? (Singangi--Ketoengau District);

Sumatra* : W. Sumatra Prov. (Indrapura), ? (Bah Lias, Baram St.);

Java* : Radja Mendala.

Figulus mecynodontus DE LISLE, 1974.

De Lisle, 1974:803; Bomans, 1986:8; Maes, 1992:39; Bomans & Bartolozzi, 1996:219.

Solomon Is. : Makira I.--10°32S-161°50E = San Cristobal I. (Bwernaniawarikiapu*).

Figulus mediocris DEYROLLE, 1874.

Deyrolle, 1874:414; Benesh, 1950; Benesh, 1960:26; Bomans, 1986:8; Bomans, 1989:13; Maes, 1992:39.

Malaya = Malacca*;

Borneo*;

Sumatra : ? (Deli);

Philippines : Mindanao I.--7°44N-125°02E.

Figulus mento ALBERS, 1883.

Figulus mento ALBERS, 1883:226.

Figulus mento subsp. foveatus KRIESCHE 1922 [Australia : Queensland : Endeavour River].

Figulus mento subsp. finschi KRIESCHE 1922 [New Guinea : Port Moresby].

Figulus mento subsp. semperi KRIESCHE 1922 [Palau I.].

Kriesche, 1922:134; Nagel, 1928:295; Benesh, 1960:26;

Figulus mento ALBERS; De Lisle, 1964

Bomans, 1984:357; Bomans, 1988:8; Maes, 1992:39; Moore & Cassis, 1992:10

Figulus mento ALBERS; Cammaerts & Bomans, 1997:13.

Note : This species is perhaps a synonym of F. sulcicollis.

Philippines : Luzon I. : Laguna

Philippines : Luzon I. : Mountain Province

Philippines : Mindanao I., 7°44 N - 125°02 E

Caroline Is. : Palau I., 7°30 N - 134°34 E

Lesser Sunda Is. : Aru I., 5°59 S - 134°30 E

New Guinea* : Papua : Amaira : Auyana

New Guinea* : Papua : Eastern Highlands : Okapa

New Guinea* : Papua : Madang Province : Baiteta primary forest, alt. 20 m, 24-VI-1993, fumigation of the crown of a Pometia pinata (Sapindaceae), col. O. Missa (1 ex. in coll. H.E. Bomans, in Natural History Museum, London).

New Guinea* : Papua : Madang Province : Madang, 5°12 S - 145°43 E

New Guinea* : Papua : Madang Province : Port Moresby, 9°28 S - 147°10 E

Australia : Queensland : Endeavour River, 15°22 S - 145°02 E.

Note : Cammaerts & Bomans (1997) tell that this species does not exist in Aru and Palau Is.

Saproxylophagous : Poinciana.

Bibliography.

DE LISLE M.O. (1964) On a collection of stag-beetles (Col., Lucanidae) from New Guinea. Papua New Guinea Scient. Soc. Trans., pp.3-13.

Figulus meridianus BOMANS, 1986.

Bomans, 1979; Bomans, 1986:12, pl.2; Maes, 1992:39; Bomans, 1994:489.

India : Kerala (Paonmudi R.*), Tamil Nadu (Madras, Jola, Anamalai Hills*, Jola--Pennumicidi*, Anamalai Hills--Kadamparai).

Figulus minor KRIESCHE, 1920.

Kriesche, 1920:114; Benesh, 1960:27; Maes, 1992:39.

Cameroon : Johann-Albrechtshohe*--4°10N-9°13E, Bipindi*;

Equatorial Guinea = Spanish Guinea : Nkolentangan*.

Figulus minutus DEYROLLE, 1874.

Deyrolle, 1874:414; Benesh, 1960:27; Maes, 1992:39.

Moluccas Is. : Banda I.*.

Figulus moluccanus DE LISLE, 1970.

De Lisle, 1970:116; Maes, 1992:39.

Moluccas Is. : Halmahera I.--0°40N-128°E = Gilolo I. (Biaur*).

Figulus morosus DE LISLE, 1974.

De Lisle, 1974:801; Bomans, 1986:8; Maes, 1992:39; Bomans & Bartolozzi, 1996:219.

Solomon Is. : Malaita I.--8°59S-160°59E (Auki*).

Figulus myrmecodianus CAMMAERTS & BOMANS, 1997.

Figulus myrmecodianus CAMMAERTS & BOMANS, 1997:7-10, figs. 1-3, 7-10.

Myrmecophilous : Phylidris sp.

In roots of Rubiaceae : Myrmecodia tuberosa.

New Guinea : Papua : Morobe Province : Patep : Lae-Bulolo road, alt. 1120 m, 2-III-1990, on woody calcareous hill, inside tuber of Myrmecodia tuberosa (Rubiaceae) hanging at 2 meters high and inhabiting by Phylidris sp. (Formicidae) (male HOLOTYPUS in Institut royal des Sciences naturelles, Bolgium).


Figulus myrmecodianus : male Holotype (adapted from Cammaerts & Bomans).

Figulus myrmecodianus : mandibles (adapted from Cammaerts & Bomans).	Figulus myrmecodianus : mentum (adapted from Cammaerts & Bomans).

Figulus myrmecodianus : left tibia I and tarsi (adapted from Cammaerts & Bomans).	Figulus myrmecodianus : male genitalia (adapted from Cammaerts & Bomans).

Bibliography.

Figulus napu KRIESCHE, 1922.

Kriesche, 1922:130; Benesh, 1960:27; Bomans, 1986:11; Maes, 1992:39.

China : Yunnan;

Vietnam : Cochinchina (Saigon*).

Figulus nitens WATERHOUSE, 1874.

Waterhouse, 1874:7; Benesh, 1960:27; Maes, 1992:39; Moore & Cassis, 1992:10.

Australia : New South Wales* (Murray--Darling Basin--32to36°S-115to148°E), Victoria.

Saproxylophagous.

Figulus nitidulus GESTRO, 1881.

Figulus nitidulus GESTRO, 1881: 335-336.

Benesh, 1960:27; Maes, 1992:39

Figulus nitidulus GESTRO; Cammaerts & Bomans, 1997:10-12, figs. 4-6, 11-14.

New Guinea : Papua : Fly River, 1876/1877, G. Doria, O. Beccari & L.M. D’Albertis Expedition (2 SYNTYPUS of Figulus nitidulus GESTRO, 1881 in Genoa Museum, Italy).

New Guinea : Papua : Northern Province : Dobodura, V/VII-1944, col. Darlington, det. B. Benesh (1 ex. in Field Museum, Chicago, USA).

New Guinea : Papua : Morobe Province : Wau, 7°20 S – 146°43 E, 1200 m, 14-IX-1961, col. J. Sedlacek (1 ex. in Bishop Museum, Hawaii).

New Guinea : Papua : Morobe Province : Wau, 7°20 S – 146°43 E, 1200 m (1 ex. in col. G. Minet, Francia).

New Guinea : Papua : East Sepik Province : Passam, 14-VIII-1989, in rotten wood of Hevea brasiliensis (Euphorbiaceae), col. R.W. Hornabrook (1 ex. in col. H.E. Bomans, in col. Natural History Museum, London).

New Guinea : Deutsch Neu Guinea (1 male in col. H.E. Bomans, in col. Natural History Museum, London).

Bismarck : Mussau I., 1°26 S – 149°38 E : Talumalaus (1 ex. in col. H.E. Bomans, in col. Natural History Museum, London).

Saproxylophagous : Euphorbiaceae : Hevea brasiliensis.


Figulus nitidulus : male Holotype (adapted from Cammaerts & Bomans).

Figulus nitidulus : mandibles (adapted from Cammaerts & Bomans).	Figulus nitidulus : mentum (adapted from Cammaerts & Bomans).

Figulus nitidulus : left tibia I and tarsi (adapted from Cammaerts & Bomans).	Figulus nitidulus : male genitalia (adapted from Cammaerts & Bomans).

Bibliography.

Figulus nubilus DIDIER, 1930.

Didier, 1930:169; Benesh, 1960:27; Maes, 1992:39; Mizunuma & Nagai, 1994:300, pl. 136.

Moluccas Is. : Ambon I.--3°39S-128°07E = Amboine I.;

Lesser Sunda Is. : Larat I.*;

New Guinea.

Figulus orientalis BOMANS, 1989.

Bomans, 1989:14, 17, 19.

Moluccas Is. : Halmahera I.--0°40N-128°E = Gilolo I.*, Damar I. = Dammer I.*, Ternate I.*--0°50N-127°15E, Bachan I. = Batchian I. = Palau Bachan I. = Batjan I.*.

Figulus orthognathus BENESH, 1950.

Benesh, 1950:54, pl.1; Benesh, 1950:51, 54-55; Benesh, 1960:27; Bomans, 1987:10; Maes, 1992:39; Sakaino & Ochi, 1993:21.

Philippines : Mindanao I.--7°44N-125°02E (Bukidnon--Santa Fe*).

Figulus papuanus GESTRO, 1881.

Gestro, 1881:336; Arrow, 1938:53; Benesh, 1955:67; Benesh, 1960:27; Bomans, 1986:12; Bomans, 1988:8; Maes, 1992:39.

Lesser Sunda Is. : Kai I.*--5°42S-132°55E;

New Guinea : Papua (Fly River*, Eastern Highlands--Kassem Pass, Madang--5°12S-145°43E--Gogol River).

Figulus parallelicanthus BOMANS, 1989.

Bomans, 1989:17-18.

Vietnam : Tonkin* (Hoah Binh*).

Figulus parvus BOMANS, 1987.

Bomans, 1987:10, pl.7; Maes, 1992:39.

Zaire : Haut-Zaire [Kibali-Ituri (Terr. Epulu--Mambasa-Mungbere*)].

Figulus philippinensis BENESH, 1955 (new name for foveatus).

Figulus foveatus BENESH, 1950:55-56 [Mountain Province : Mt. Pulay, Laguna : Mt. Makilling] (pre occupied name).

Figulus philippinensis BENESH, 1955:64 (new name for foveatus).

Benesh, 1960:27; Bomans, 1987:10; Maes, 1992:40.

Figulus philippinensis BENESH; Bomans & Benoit, 2005:291.

Ø Philippines : Luzon I. : Mountain Province : Mt. Pulay

Ø Philippines : Luzon I. : Laguna : Mt. Makilling.

Ø Philippines : Mindanao I. : Santa Fé : Bukidnon, 2300 Ft., 15-V-1935, ex coll. Van Dyke (TYPUS in California Academy of Sciences, San Francisco, USA).

Bibliography :

Figulus piceus BOMANS, 1989.

Bomans, 1989:15.

Thailand = Siam*;

Vietnam : Saigon *.

Figulus planifrons BOMANS, 1987.

Bomans, 1987:6, pl.5; Maes, 1992:40.

Sumatra. : N. Sumatra Prov. (Kora Kora--Dolok Merangir*), ? (Pagar Marbau*).

Figulus popei BOMANS, 1986.

Bomans, 1986:8; Maes, 1992:40; Bomans & Bartolozzi, 1996:219.

Solomon Is. : Makira I.--10°32S-161°50E = San Cristobal I. (Kira-Kira*).

Figulus porrectus BOMANS, 1989.

Bomans, 1989:14, 17, 18.

Sumatra N. Sumatra Prov. (Sindar Raja*, Dolok Meranjir*, Stabat-Langkat*).

Figulus powelli BOMANS, 1986.

Bomans, 1986:12, pl.2; Maes, 1992:40.

Borneo : Brunei*.

Figulus praecipuus BOMANS, 1987.

Bomans, 1987:13, pl.8; Maes, 1992:40.

N. China*.

Figulus procericornis BOMANS, 1987.

Figulus procericornis BOMANS, 1987:10, pl.7 (Philippines)

Figulus procericornis BOMANS; Maes, 1992:40.

Figulus procericornis BOMANS; Bomans & Benoit, 2005:292.

Ø Philippines (HOLOTYPUS in coll. H. Bomans in Natural History Museum, London).

Bibliography :

Figulus procerus HELLER, 1898.

Heller, 1898:21, pl.3; Benesh, 1960:27; Maes, 1992:40; Mizunuma & Nagai, 1994:299, pl. 136.

Sulawesi = Celebes : Mt. Bonthain*, Bua-Kraeng*, Mt. Lompobatang.

Figulus punctatus WATERHOUSE, 1873.

Waterhouse, 1873:278; Lewis, 1883:339; Miwa, 1934:320, pl.4; Benesh, 1960:27; Nomura, 1960:44; Nomura, 1966:69; Bomans, 1987:9; Fujita & Ichikawa, 1986:24; Isida & Fujioka, 1988:2; Abe, Kudoh, Ichikawa & Saitoh, 1992:218; Araya, Obuchi & Ochi, 1992:32; Maes, 1992:40; Sakaino, Yu & Chu, 1994:122; Mizunuma & Nagai, 1994:299, pl. 136; Araya & Obuchi, 1994:29, 30; Araya, 1996:21.

Japan : Honshu I. [Izu Is. (Kozushima I., Miyakejima I.--34°04N-139°30E, Mikurajima I., Hachijojima I.--33°07N-139°48E), Wakayama Pref. (Taiji), Hyogo Pref. (Nushima I.), Yamaguchi Pref.], Shikoku I. [Kochi Pref. (Okinoshima I.--32°43N-132°34E)], Kyushu I. [Fukuoka Pref. (Kitakyushu City--33°52N-130°47E--Shiroshima I.), Saga Pref. (Takashima I.--33°27N-129°47E--Higashi-matsuura Co.), Nagasaki Pref. (Nagasaki--32°47N-129°54E, Tsushima I.--34°20N-129°20E), Goto Is. (Hiradoshima I.--33°16N-129°28E, Kabeshima I.--32°46N-129°01E), Danjogunto Is., Kumamoto Pref.-- Amakusa Is.--32°23N-130°04E (Shimojima I.)], Kagoshima Pref. [Yakushima--30°22N-130°31E, Kuchinoerabujima I., Takarajima Is. = Tokara Is. (Kuchinoshima I., Nakanoshima I., Akusekijima I.), Amami-oshima I., Tokunoshima I.], Okinawa Pref. [Okinawa I.--26°30N-128°E = Okinawahonto I., Ishigakijima I.--24°24N-124°10E];

Taiwan.

Saproxylophagous : Ulmaceae : Celtis.

Figulus punctifrons BOMANS, 1987.

Bomans, 1987:9, pl.6; Maes, 1992:40.

Borneo*.

Figulus quasisimilis BOMANS, 1989.

Bomans, 1989:16.

Borneo : Sarawak*.

Figulus regularis WESTWOOD, 1834.

Figulus pacificus DEJEAN 1837 [nomen nudum] [Nov. Holland.].

Figulus australicus THOMSON 1862 [Australia].

Westwood, 1834:120; Dejean, 1837:194; Westwood, 1838:263; Burmeister, 1847:437; Thomson, 1862:432; Parry, 1863:450; Gestro, 1881:334; Heyne & Taschenberg, 1908:57; Boileau, 1913:265; Mjoberg, 1916:13; Nagel, 1928:295; Benesh, 1960:27; Holloway, 1960:343; Holloway, 1969:974; Klausnitzer, 1982:16; Hawkeswood, 1985:200; Bomans, 1986:9; Hawkeswood, 1988:270-272; Maes, 1992:40; Moore & Cassis, 1992:10; Mizunuma & Nagai, 1994:300, pl. 136.

Lesser Sunda Is. : Aru I.--5°59S-134°30E;

New Guinea : Soron, Papua (Fly River), Katau, Yule I.;

Australia = New Holland* : Northern Territory, Queensland (Yarrabah--Mission Station--16°55S-145°52E, Mt. Bellenden Ker--17°16S-145°24E, Colosseum--24°24S-151°37E, Dalby--27°11S-151°15E, Brisbane--27°28S--153°01E, Cape York--33°15S-150°08E, Christmas Creek), Torres Straits Is. (Tawan I.-- 22°10S-152°47E), New South Wales (Murray--Darling Basin--32to36°S-115to148°E), South Australia (Adelaide--34°56S-138°36E).

Saproxylophagous :

Myrtaceae : Eucalyptus.

Xanthorrhoeaceae : Xanthorrhoea.

Figulus robustus BOMANS, 1986.

Bomans, 1986:8; Maes, 1992:40; Bomans & Bartolozzi, 1996:219.

Solomon Is. : Guadalcanal I.*--9°37S-160°02E, Honiara I.*, Kukum I.*.

Figulus rossi GAHAN, 1900.

Gahan, 1900:96; Benesh, 1960:27; Maes, 1992:40; Moore & Cassis, 1992:10.

Australia : Christmas I.*--10°S-105°E;

Cocos Is. (Keeling I.--12°03S-96°54E).

Figulus rubripes BOMANS, 1987.

Bomans, 1987:5, pl.5; Maes, 1992:40.

New Guinea : Papua [Redscar Bay*, Morobe--7°42S-147°34E (Gurakor*, Kaiapit*)].

Figulus rugosus DEYROLLE, 1874.

Deyrolle, 1874:413; Benesh, 1960:27; Maes, 1992:40.

Borneo*;

Sumatra.

Figulus samoanus KRIESCHE, 1920.

Kriesche, 1920:106; Benesh, 1960:27; Maes, 1992:40.

Samoa Is.* : Upolu I.--13°55S-171°44W.

Figulus sarawakensis BOMANS, 1987.

Bomans, 1987:9, pl.6; Bomans, 1989:13; Maes, 1992:40.

Borneo : Sarawak (Kapit*).

Figulus scaritiformis PARRY, 1862.

Figulus manicatus DIDIER & SEGUY 1953 [Viet Nam : Tonkin : Chapa].

Parry, 1862:113; Parry, 1864:64; Gestro, 1881:339; Gravely, 1915:431; Didier & Séguy, 1953:108; Benesh, 1960:26,27; Bomans, 1986:12; Maes, 1992:40.

Viet Nam : Tonkin (Chapa);

Malaya = Malacca* : Johor;

Borneo : Sarawak;

New Guinea.

Figulus schismonotus (DE LISLE, 1975) [Cardanus].

De Lisle, 1975:273; Bomans, 1986:292; Maes, 1992:40.

New Guinea : Papua (Kikori*).

Figulus scotti BENESH, 1955.

Benesh, 1955:61; Benesh, 1960:154; Vinson, 1967:321; Maes, 1992:40.

Mauritius I.*--20°17S-57°34E.

Figulus serratus ARROW, 1938.

Arrow, 1938:60, pl.4; Benesh, 1960:28; Bomans, 1986:12; Maes, 1992:40; Mizunuma & Nagai, 1994:301, pl. 136.

Borneo : Natuna Is. (Bunguran Besar I.--3°56N-108°15E), Sarawak (Retuh*, Tabehang*).

Figulus seychellensis SCOTT, 1912.

Scott, 1912:222; Benesh, 1960:28; Maes, 1992:40.

Seychelles Is. : Silhouette I.*--4°28S-55°14E.

Figulus similis BOMANS, 1987.

Bomans, 1987:5, pl.5; Maes, 1992:41.

Moluccas Is. : Waigeo I.--0°07S-130°50E = Waigiou I.*;

New Guinea : Irian Jaya*.

Figulus spinosus BOMANS, 1986.

Bomans, 1986:8; Maes, 1992:41.

Solomon Is. : Guadalcanal I.*--9°37S-160°02E (Mt. Austen*), Popomanasiu I., Kukum I.*, Honiara I.*.

Figulus splendens BOMANS, 1986.

Bomans, 1986:11; Bomans, 1988:8; Maes, 1992:41.

New Guinea : Papua (Fly River--Kiunga*).

Figulus striatus (OLIVIER, 1789) [Lucanus].

Figulus striatus var. braueri KRIESCHE 1922 [Seychelles Is. : Mamelle I.].

Olivier, 1789:19-20, pl.4; Fabricius, 1801:253; W.S. Macleay, 1819:110; Hope, 1837:79; Dejean, 1837:194; Westwood, 1838:262; Castelnau, 1840:175, pl.17; Blanchard, 1845:268, pl.8; Burmeister, 1847:438; Coquerell, 1866:31; Kolbe, 1910:22; Sharp & Muir, 1912:575; Scott, 1912:221; Mjoberg, 1916:14; Kriesche, 1922:130; Paulian, 1957:95; Benesh, 1960:28; Vinson, 1967:321; Fujita & Ichikawa, 1986:29; Maes, 1992:41; Moore & Cassis, 1992:10-11; Mizunuma & Nagai, 1994:300, pl. 136.

Madagascar;

Reunion I., 21°06S - 55°31E = Bourbon I.* : Brulé de Mare-Longue

Mauritius I., 20°17S - 57°34E = Isle of France

Aldabra I., 9°23 S - 46°20 E

Seychelles Is. : Silhouette I., 4°29 S – 55°14 E : Gratte Fesse, 400 m, 3-VIII-1998, in rotten Paraserianthes falcataria wood, col. J. Gerlach (2 ex in Nature Protection Trust of Seychelles, ref. NPTS Hc 1998 - 114 & 115).

Seychelles Is. : Mamelle I., 4°29S - 55°31E.

[No Australia].

Saproxylophagous : Paraserianthes falcataria.

Figulus subcastaneus WESTWOOD, 1838.

Figulus haedulus DEJEAN 1837 [nomen nudum] [Java].

Dejean, 1837:194; Westwood, 1838:263; Burmeister, 1847:438; Oberthur & Houlbert, 1914:225; Benesh, 1960:28; Maes, 1992:41.

Borneo : Sarawak (Baram St.);

Java : Mt. Tengger.

Figulus sublaevis (BEAUVOIS, 1805).

Beauvois, 1805:3, pl.1; Schonherr, 1817:331; Klug, 1833:85; Westwood, 1834:120, pl.7; Dejean, 1937:194; Westwood, 1838:261-262; Erichson, 1839:336; Burmeister, 1847:436; Parry, 1862:113; Thomson, 1862:433; Parry, 1864:7; Parry, 1864:64, 99; Gemminger & Harold, 1868:965; Gerst., 1871:46; Parry, 1873:343; Parry, 1875:22; Albers, 1884:173; Albers, 1885:248, 284; Heyne & Taschenberg, 1908:57, pl.8; Boileau, 1909:338; Peringuey, 1909:548; Kriesche, 1920:112-114; Didier & Séguy, 1953:57; Brinck, 1956:314; Benesh, 1960:28; Van Waerebeke, 1970:6, 10; Gomes Alves, 1973:30; Klausnitzer, 1982:36; Bomans, 1992:251-252; Maes, 1992:53; Maes, 1992:41; Mizunuma & Nagai, 1994:300, pl. 136.

Figulus sublaevis ssp. decipiens ALBERS, 1884 [West Africa].

Principe I.--1°37N-7°23E;

Sao Tomé I.--0°13N6°31E : Roca Diego Vaz, Roca Monte Cafe, Binda.

Figulus sublaevis ssp. sublaevis (BEAUVOIS, 1805) [Lucanus] [West Africa].

Figulus anthracinus KLUG 1833 [Madagascar].

Figulus ebenus WESTWOOD 1834 [Madagascar].

Figulus ovis DEJEAN 1837 [nomen nudum] [Senegal].

Figulus nigritus WESTWOOD 1838 [Senegal].

Figulus monilifer PARRY 1862 [New Zealand].

Figulus vulneratus THOMSON 1862 [Madagascar].

Figulus cernensis (HOPE) GEMMINGER & HAROLD 1868 [Madagascar].

Figulus lettowi KRIESCHE 1920 [Tanzania : Daressalam, Uhene--Iringa, Langenburg, Unyika].

Figulus nossibenus KRIESCHE 1920 [Madagascar : Nossi Be I.].

Senegal*;

Guinea : Mt. Nimba Park, 7°37 N - 8°25 W : Yalanzou

Ivory Coast : Bingerville;

Togo : Bismarckburg--8°11N-0°39E;

Benin : 100 km de Cotonou : Forêt de la Lama, 26-V-2001, col. S. Attignon & T. Lachat (Séverin Tchiboso, in litteris).

Cameroon : Johann-Albrechtshohe--4°10N-9°13E, Moliwe near Victoria;

Principe I.--1°37N-7°23E : Roca Sundy;

Zaire : Shaba [Upemba Park (Kanonga), Upemba Lake (Mabwe), Kanakakazi River (Masombwe--9°05S-27°12E), Kaziba--7°09S-27°01E];

Angola : Dundo;

Tanzania : Iringa [Iringa--7°44S-35°38E (Uhene), Langenburg--9°36S-34°15E], Pwani [Dar Es Salaam], Ruvuma [Unyika];

Zambia : Kankunda--10°19S-30°03E;

Mozambique : Garuzo, Caia--17°45S-35°18E, Beira--19°44S-34°52E;

Comoros Is. : Mayotte I.--12°47S-45°08E;

Madagascar : Mananjari, Fenerivo, Amboasary--Moramanga Forest, Nosy Be I. --13°17S-48°14E.

Parasite : Nematoda : Thelastoma toxi, Thelastoma figuli.

Note : Kriesche divide this species into the following ssp. : ssp. nigrita from Senegal; ssp. sublaevis from Togo to Cameroon; ssp. lettowi from Tanzania; ssp. nossibenus from Nosy Be I. and ssp. anthracinus from Madagascar.

Figulus sulcicollis HOPE & WESTWOOD, 1845.

Figulus sulcicollis HOPE & WESTWOOD, 1845:26.

Figulus ater DEYROLLE, 1874:412.

Gestro, 1881:337; Benesh, 1960:24, 29; Bomans, 1986:9; Bomans, 1988:8; Maes, 1992:36, 41; Moore & Cassis, 1992:11

Figulus sulcicollis HOPE & WESTWOOD; Cammaerts & Bomans, 1997:12-13, 15-19, figs. 15-29.

Moluccas Is. : Bachan I. = Batchian I.

Moluccas Is. : Misool I., 1°54 S - 130°05 E = Mysole I.*

Moluccas Is. : Andai I.

New Guinea : Papua : Madang Province : Karkar I., 4°38 S – 145°59 E : Kurumlang : south of Kulkul plantation, alt. 50-100 m, 18-XII-1989, under bark of a dead tree, in rainforest very deteriored, volcanic soil (1 ex. in col. H.E. Boman, in Natural History Museum, London).

New Guinea : Papua : Laing I., alt. 1 m, 23-I-1990, coastal forest, under bark of dead Terminalia catappa (Combretaceae) (1 adult, 5 larvae + 1 group of 2 adults and 4 larvae, in col. H.E. Boman, in Natural History Museum, London).

New Guinea : Papua : Laing I., alt. 1 m, 1-II-1990, coastal forest, under bark of dead Terminalia catappa (Combretaceae) (1 group of 3 adults and 9 larvae + 1 group of 2 adults and 4 larvae, in col. H.E. Boman, in Natural History Museum, London).

New Guinea : Papua : Laing I., alt. 1 m, 19-IV-1990, coastal forest, inside trunk of live Excoecaria agallocha (Euphorbiaceae) (1 ex. in col. H.E. Boman, in Natural History Museum, London).

New Guinea : Papua : Morobe Province : Aseki, 7°21 S – 146°12 E, alt. 1600 m, on a trunk of Leucaena leucophylla (Fabaceae), in a coffee plantation on calcareous soil (1 ex. in col. H.E. Boman, in Natural History Museum, London).

Australia : Torres Straits Is.

Australia : Northern Territory : Melville I., 11°34 S - 130°56 E

Australia : Northern Territory : Port Essington*

Australia : Queensland : Cairns, 16°54 S - 145°46 E

Australia : Queensland : Innisfail, 17°32 S - 146°01 E

Saproxylophagous : Combretaceae : Terminalia catappa.

Xylophagous : Euphorbiaceae : Excoecaria agallocha.

Larvae of Figulus sulcicollis : habitus and antena (adapted from Cammaerts and Bomans).

Larvae of Figulus sulcicollis : epipharynx and mandibles in ventral view (adapted from Cammaerts and Bomans).

Larvae of Figulus sulcicollis : hypopharynx and mandibles in internal view (adapted from Cammaerts and Bomans).

Larvae of Figulus sulcicollis : leg II and pars stridens (adapted from Cammaerts and Bomans).

Larvae of Figulus sulcicollis : plectrum (adapted from Cammaerts and Bomans).

Larvae of Figulus sulcicollis : leg III and apex of leg III (adapted from Cammaerts and Bomans).

Larvae of Figulus sulcicollis : anal sclerite and microscopical view of the claw of leg III (adapted from Cammaerts and Bomans).

Bibliography.

Figulus tambourinensis MJOBERG, 1916.

Mjoberg, 1916:13; Nagel, 1939:1; Benesh, 1960:29; Maes, 1992:41; Moore & Cassis, 1992:11.

Australia : Queensland (Mt. Tambourine*--27°55S-153°10E).

Saproxylophagous.

Figulus tapinus BOMANS, 1986.

Bomans, 1986:7; Maes, 1992:41.

India : Uttar Pradesh (Dehra Dun*).

[Figulus tibialis (ALBERS) NAGEL, 1924 nomen nudum].

Nagel, 1924; Nagel, 1926; Benesh, 1960:152; Maes, 1992:41.

Figulus trilobus WESTWOOD, 1838 [Nigidius].

Westwood, 1838:263; Burmeister, 1847:439; Parry, 1873:343; Parry, 1875:15; Benesh, 1943:46; Benesh, 1960:29; Maes, 1992:41; Moore & Cassis, 1992:11.

Australia = New Holland* : New South Wales (Wauchope--31°28S-152°44E, Murray--Darling Basin--32to36°S-115to148°E).

Saproxylophagous.

Figulus tubericollis BOMANS, 1989.

Bomans, 1989:16, 17.

Vietnam : Annam (Phuc-Son*), Cochinchina*.

Figulus tumidimentum BOMANS, 1989.

Bomans, 1989:17, 18-19.

India : Tamil Nadu : (Trichinopoly*).

Figulus venustus BOMANS, 1989.

Bomans, 1989:16, 17.

South Korea : Soul--37°29N-127°00E = Seoul*.

Figulus vinsoni BENESH, 1955.

Benesh, 1955:62; Benesh, 1960:154; Vinson, 1967:321; Maes, 1992:41.

Mauritius I.*--20°17S-57°34E.

Figulus weinreichi BOMANS, 1986.

Bomans, 1986:15, pl.2; Maes, 1992:41.

New Guinea : Papua [Morobe--7°42S-147°34E (Kaiapit*)], D.New Guinea*.

Figulus wittmeri BOMANS & LACROIX, 1989.

Bomans & Lacroix, 1989:24; Bartolozzi & Sforzi, 1994:114.

Nepal : Biratnagar*.

Figulus yujii FUJITA, 1994.

Fujita, 1994:15; Araya, 1996:21.

Japan : Tokyo--Volcano Is. (Naka-iwojima I.--24°45N-141°20E).

Figulus yunnanensis NAGEL, 1928.

Nagel, 1928:295; Benesh, 1960:29; Bomans, 1986:11; Bomans, 1989:9; Maes, 1992:41.

China : Yunnan (Tche Ping Tcheou* - 30°16 N - 102°24 E), Fujian (Kuatun);

Vietnam: Tonkin.

HOME	LUCANIDAE
GENERA