Calcodes WESTWOOD, 1834

(G. = Lucanus aeratus)

= Chalcodes BURMEISTER, 1847 (G. = Lucanus aeratus)

= Odontotyranus SEGUY, 1955 (G. = Odontolabis striatus)


Calcodes aeratus (HOPE, 1834).

Lucanus aeratus HOPE, 1834:118 [Tenasserim] [Anoplocnemus, Odontolabis, Chalcodes] [aerata].

Hope, 1835:99, pl.14; Castelnau, 1840:172; Hope & Westwood, 1845:16; Burmeister, 1847:361; Westwood, 1848:22, pl.10; Parry, 1864:pl.7; Parry, 1870:74; Leuthner, 1885:473, pl.97; Zang, 1905:214; Heyne & Taschenberg, 1908:52; Gravely, 1915:416; Arrow, 1937:241, pl.3; Arrow, 1950:209, pl.20; Arrow, 1951:pl.15; Didier & Séguy, 1952:pl.41; Didier & Séguy, 1953:91; Benesh, 1960:121; Weinreich, 1972; Lacroix, 1978:262; Lacroix, Ratti & Taroni, 1983:2; Weng-Yew Tung, 1983:49, pl.4; Lacroix, 1984:34-35, pl.1; Kikuta, 1986:135; Nagai, 1986:3, 10; Bartolozzi & Bomans, 1990:438; Bomans, 1991:145; Maes, 1992:22.

Calcodes aeratus (HOPE); Mizunuma & Nagai, 1994:16-17, fig. 38(a-f), p. 230, pl. 38.

Araya, 1996:17.

Myanmar = Burma : Tenasserim* State.

Thailand : Surat Thani.

Malaya = Malacca : Kedah.

Malaya : Pinang : Pinang I., 5°22N-100°12E (= Prince of Wales I.).

Malaya : Perak : Taiping, 4°51N-100°41E.

Malaya : Perak : Kampong Sahom.

Malaya : Perak : Cameron Highlands : Taphh.

Malaya : Pahang : Cameron Highlands.

Malaya : Pahang : Tengah-Geb.

Malaya : Pahang : Fraser's Hills.

Malaya : Singapore : Chapais.

Borneo : Natuna Is. : Bunguran Besar I., 3°56N-108°15E.

Borneo : W. Kalimantan : Sambas.

Sumatra : W. Sumatra Prov. : Sinkep I.

Sumatra : S. Sumatra Prov. : Palembang.

Sumatra : Siboga.

Java.

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Calcodes baderi (BOMANS & BARTOLOZZI, 1990) [Chalcodes, Odontolabis].

Bomans & Bartolozzi, 1990:433; Maes, 1992:22; Mizunuma & Nagai, 1994:16, 231, pl. 39; Ipsen, 1995:1; Ikeda, 1996:4.

Sulawesi = Celebes* : Palopo--2°55S-120°06E (Pulu Pulu).

 

Calcodes carinatus (LINNAEUS, 1758) [Scarabaeus, Odontolabis, Chalcodes] [carinata] [India].

= Lucanus bengalensis TENNENT 1859.

= Odontolabis cingalensis PARRY 1864 [Ins. Taprobana].

= Odontolabis intermedia DEYROLLE 1864 [Ceylon].

= Odontolabis nigrita DEYROLLE 1864 [Chalcodes] [nigritus] [Ceylon].

= Chalcodes srilankaensis BOMANS 1974 [Ceylan].

Linnaeus, 1758:353; Linnaeus, 1764:34; Linnaeus (ed. Gmelin), 1790:1590; Herbst, 1790:315; Muller, 1776:97; Thunberg, 1806:193, pl.12; Schonherr, 1817:323; Tennent, 1859:276; Deyrolle, 1864:314; Parry, 1864:16, 77, pl.10; Sharp, 1884:18; Leuthner, 1885:474, pl.97; Heyne & Taschenberg, 1908:52, pl.7; Boileau, 1913:243; Gravely, 1915:416; Arrow, 1937:241, pl.1; Arrow, 1950:207, pl.20; Arrow, 1951:pl.15; Didier & Séguy, 1952:pl.40; Didier & Séguy, 1953:91; Benesh, 1960:114; Bomans, 1974:181; Lacroix, 1978:258; Numhauser, 1981:74; Lacroix, 1984:39-43,             pl.3-4; Bartolozzi & Bomans, 1988:88-89; Bartolozzi & Bomans, 1990:433; Maes, 1992:22; Mizunuma & Nagai, 1994:229, pl. 38.

Sri Lanka = Ceylon : Western Prov. [Colombo Distr. (Labugama), Kalutara Distr. (Agalawatta)], Sabaragamuwa Prov. [Ratnapura Distr. (Rakwana, Bulutota, Gilimale, Sinharaja Jungle 3 mi S Weddagala)], Central Prov. [Kandy Distr. (Kandy, Karabagala--Willomalai Jungle), Nuwara Eliya Distr. (Bogawantalawa, Maskeliya, Lindula)], Uva Prov. [Badulla Distr. (Ohiya--W. Haputale, Haldumulla)], Southern Prov. [Galle Distr. (Udugama--Kaneliya, Kaneliya Jungle, Sinharaja Jungle, Galle)], ? [Taprobana, Mousakande].

 

Calcodes cephalotes (LEUTHNER, 1885) [Odontolabis, Chalcodes].

= Odontolabis striata WESTWOOD (nec DEYROLLE) 1874 (en parte).

Westwood, 1874; Leuthner, 1885:478, pl.97; Benesh, 1960:115; De Lisle, 1976:276; Klopp, 1977:60; Lacroix, 1978:259; Lacroix, 1984:45-47, pl.5; Nagai, 1986:8; Bartolozzi & Bomans, 1990:438; Maes, 1992:22.

Borneo*.

 

Calcodes cupreiventris (MOLLENKAMP, 1900).

Odontolabis cupreiventris MOLLENKAMP, 1900:45 [Borneo].

Mollenkamp, 1903:348; Didier & Séguy, 1953:98; Benesh, 1960:121; Lacroix, 1978:262; Lacroix, 1984:36-37, 46, pl.5; Nagai, 1986:10; Bartolozzi & Bomans, 1990:438; Maes, 1992:22.

Calcodes cupreiventris (MOLLENKAMP); Mizunuma & Nagai, 1994:17, fig. 39(a-b, f), p. 230, pl. 38.

Araya, 1996:17.

Borneo : Sarawak : Mt. Serapi near Kuching.

Borneo : W. Kalimantan : Mt. Bawang.

Sumatra : Jambi Prov. : O.K.Siantar-Simpang Raja.

Sumatra : Sibolangit.

Sumatra : Langkat-Balei Gadjah.

Sumatra : Deli.

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Calcodes invitabilis (MIZUNUMA, 1994).

Odontolabis invitabilis MIZUNUMA in MIZUNUMA & NAGAI, 1994:16-17, fig. 37(a-c). p. 229, 230, pl. 139, fig. 183(1-2) [Berastagi].

Sumatra : N. Sumatra Prov. : near Berastagi, IV-1993 (male HOLOTYPUS in Kanagawa Prefectural Museum, Japan).     

Sumatra : N. Sumatra Prov. : near Berastagi, V/VII-1993 (3 males PARATYPUS in col. T. Mizunuma, Japan) (1 male PARATYPUS in col. H. Fujita, Japan).  

Sumatra : N. Sumatra Prov. : near Berastagi (1 male PARATYPUS in col. H. Yasuoka, Japan).  

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Calcodes kazuhisai (MIZUNUMA, 1994).

Odontolabis kazuhisai MIZUNUMA in MIZUNUMA & NAGAI, 1994:16-17, fig. 36(a-b, f), pl. 38, fig. 184(1-3) [ Mt. Singgalang].

Kawano, 1997:454.

Sumatra : W. Sumatra Prov. : Padang : Padangpanjang : Mt. Singgalang, 7-VIII-92, leg. Kazuhisa Fujita (male HOLOTYPUS in Kanagawa Prefectural Museum, Japan) (1 male PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : W. Sumatra Prov. : Padang : Padangpanjang : Mt. Singgalang, 28-VII-92, leg. Kazuhisa Fujita (female ALLOTYPUS in Kanagawa Prefectural Museum, Japan).

Sumatra : W. Sumatra Prov. : Padang : Padangpanjang : Mt. Singgalang, II/III-93 (16 males PARATYPUS in col. T. Mizunuma, Japan).

Sumatra : W. Sumatra Prov. : Padang : Padangpanjang : Mt. Singgalang, III-93 (4 males PARATYPUS, 1 female PARATYPUS in col. H. Fujita, Japan).

Sumatra : W. Sumatra Prov. : Padang : Padangpanjang : Mt. Singgalang, V-93 (30 males PARATYPUS, 30 females PARATYPUS in col. T. Mizunuma, Japan) (2 females PARATYPUS in col. T. Mizukami, Japan).

Sumatra : W. Sumatra Prov. : Padang : Padangpanjang : Mt. Singgalang (6 males PARATYPUS, 2 females PARATYPUS in col. H. Yasuoka, Japan) (4 males PARATYPUS, 2 females PARATYPUS in col. M. Suzumura, Japan).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Calcodes lecourti (BOMANS, 1997) [Chalcodes].

Bomans, 1997 : 29-30.

Sulawesi = Celebes : Palopo--2°55S-120°06E (Rantepao--Pulu-Pulu*).

 

Calcodes martinii (IPSEN, 1995) [Chalcodes].

Ipsen, 1995:1-5.

Sulawesi : Palopo--2°55S-120°06E (Rantepao*).

 

Calcodes nagaii (IKEDA, 1996) [Odontolabis].

Ikeda, 1996:3-6.

Sulawesi = Celebes : Palopo--2°55S-120°06E (Pulu Pulu*).

 

Calcodes nishiyamai (MIZUNUMA, 1994).

Odontolabis nishiyamai MIZUNUMA in MIZUNUMA & NAGAI, 1994:16-17, fig. 40(a-h), pp. 231-232, pl. 39, fig. 191(1-14) [Ende].

Ikeda, 1996:3-4.

Lesser Sunda Is. : Flores I. (8°37S-121°19E) : near Ende, XI/XII-1991 (male HOLOTYPUS, 3 males PARATYPUS in Kanagawa Prefectural Museum, Japan) (female ALLOTYPUS, 84 males PARATYPUS in col. T. Mizunuma, Japan) (4 males PARATYPUS in col. M. Suzumurai, Japan) (10 males PARATYPUS in col. H. Fujita, Japan) (6 males PARATYPUS, 3 females PARATYPUS in col. M. Baba, Japan) (3 males PARATYPUS in col. T. Mizukami, Japan).

Bibliography.

MIZUNUMA T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156 pls.

 

Calcodes ritsemae (BOILEAU, 1899) [Odontolabis, Chalcodes] [ritsemai].

= Odontolabis liewi DE LISLE 1976 [Malaya : Maxwell's Hill].

= Odontolabis albolateralis KLOPP 1977 [Malaya : Perak : Taiping].

Boileau, 1899:42; Benesh, 1960:119; De Lisle, 1976:274-277; Klopp, 1977:60; Lacroix, 1984:48-49, pl.6; Bartolozzi & Bomans, 1990:438; Maes, 1992:23; Mizunuma & Nagai, 1994:229, pl. 38.

Malaya = Malacca* : Perak (Taiping--4°51N-100°41E, Taiping--Maxwell's Hill), Pahang (Cameron Highlands);

Sumatra : N. Sumatra Prov. (Dolok Merangir).

 

Calcodes rorekaotimbuensis (NAGAI & ISOGAI, 1987) [Odontolabis, Chalcodes].

Nagai & Isogai, 1987:9-11; Bartolozzi & Bomans, 1990:436; Maes, 1992:23; Mizunuma & Nagai, 1994:16, 231, pl. 39; Ipsen, 1995:1; Ikeda, 1996:3-4.

Sulawesi = Celebes : Mt. Rorekaotimbu--Puncak Dingin*.

 

Calcodes rubens (LACROIX, 1978) [Chalcodes].

Lacroix, 1978:264; Lacroix, 1984:37-38, 42, pl.4; Bartolozzi & Bomans, 1990:438; Maes, 1992:23; Mizunuma & Nagai, 1994:230.

Malaya : Pahang (Kuala-Lipis*);

Borneo : Sarawak (Mt. Serapi near Kuching), W. Kalimantan (Mt. Bawang near Singkawang).

 

Calcodes sarasinorum (HELLER, 1898) [Odontolabis, Chalcodes].

Heller, 1898:21, pl.3; Benesh, 1960:119; De Lisle, 1976:277; Lacroix, 1984:49-50, pl.6; Nagai & Isogai, 1987:9-10; Bartolozzi & Bomans, 1990:438Maes, 1992:27; Mizunuma & Nagai, 1994:16, 231, pl. 39; Ikeda, 1996:4.

S. Sulawesi = Celebes : Mt. Bonthain*

S. Sulawesi : Sinjai

S. Sulawesi : Mt. Bawakaraeng, I-2001, leg. M. Djufri (2 males, 1 female in col. Ronald Cave, USA).

 

Calcodes striatus (DEYROLLE, 1864) [Odontolabis, Chalcodes, Odontotyranus] [striata].

Deyrolle, 1864; Westwood, 1874:360, pl.3; Boileau, 1913:241; Didier & Séguy, 1952:pl.39; Séguy, 1955:35; Benesh, 1960:120; De Lisle, 1976:276; Klopp, 1977:60; Klausnitzer, 1982:29; Lacroix, 1984:44-45, pl.5; Nagai, 1986:3, 8-10; Bartolozzi & Bomans, 1990:438; Maes, 1992:23; Mizunuma & Nagai, 1994:229, pl. 38, 139; Araya, 1994:4.

Malaya = Malacca* : Johor (Kuantan--3°50N-103°18E);

Borneo : Sarawak (Mt. Serapi near Kuching), Sabah (Telupid, Crocker Range, Sandakan), W. Kalimantan (Pontianak, Putusibau, Karimata I.), ? (Gunong-Gadin-Lundu);

Sumatra : W. Sumatra Prov. (Mt. Singgalang), Belitung I.;

Java.

Note : Perhaps it's safe to use the subgenus Odontotyranus SEGUY for the Chalcodes with spinose jugae, like C. striatus, C. ritsemae and C. cephalotes.

 Bibliography.

ARAYA K. (2000) An account of a visit to European Museums. 2. The lucanid specimens in the van Roon and Siebold collections deposited in the Rijksmuseum van Natuurlijke Historie, Leiden. Gekkan-Mushi, 350:4-16.

 

Calcodes taronii (BOMANS & BARTOLOZZI, 1990) [Chalcodes, Odontolabis] [tarorii].

Bomans & Bartolozzi, 1990:436; Maes, 1992:23; Mizunuma & Nagai, 1994:16, 231, pl. 39; Ipsen, 1995:1; Ikeda, 1996:3-4.

S. Sulawesi = Celebes* : Palopo--2°55S-120°06E (Puncak*, Pulpul*, Pulu Pulu), Bantaeng*.


HOME

LUCANIDAE

GENERA