Lucanus (Lucanus) SCOPOLI, 1763
(G. =
Scarabaeus cervus)
= Hexaphyllus MULSANT 1839 (G. = Hexaphyllus
pontbrianti)
Lucanus
(Lucanus) angusticornis DIDIER, 1925.
Lucanus
angusticornis (Boileau in litt.) DIDIER, 1925:219 [Laos : Xieng-Khouang].
Lucanus angusticornis DIDIER; Didier, 1926:144
Lucanus angusticornis DIDIER; Didier & Séguy, 1953:81
Lucanus angusticornis DIDIER; Benesh, 1960:131
Lucanus angusticornis DIDIER; Lacroix, 1972:50
Lucanus angusticornis DIDIER; Kurosawa, 1978:143
Lucanus angusticornis DIDIER; Sakaino, 1981:23
Lucanus angusticornis DIDIER; Maes, 1992:15
Lucanus angusticornis DIDIER; Kawano, 1994:9
Lucanus angusticornis DIDIER; Mizunuma & Nagai, 1994:216, pl.15
Lucanus angusticornis DIDIER; Kawano, 1997:454
Lucanus angusticornis DIDIER; Kawano, 1997:25.
Lucanus angusticornis DIDIER; Araya, 2001:10, figs.1-5.
·
Laos : Xieng-Khouang, 18-VI-1919, ex. coll. Boileau (male HOLOTYPE of Lucanus
angusticornis [Boileau in litteris] DIDIER, 1925 en Museum National
d’Histoire Naturelle, Paris).
·
Vietnam : Tonkin : Tam Dao near Vinh Yen at NE Hanoi, leg. H. Perrot
(female TYPE of Lucanus angusticornis DIDIER, 1925 by LACROIX, 1969,
published in 1972 en Museum National d’Histoire Naturelle, Paris).
|
|
|
|
Male HOLOTYPE of Lucanus angusticornis, from
Laos (adapted from Araya, 2001). |
Female TYPUS of Lucanus angusticornis, from
Viet Nam (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Lucanus) bruanti LACROIX & BOMANS, 1973.
Lucanus
bruanti LACROIX & BOMANS, 1973:138
Lucanus
bruanti LACROIX & BOMANS; Mizunuma & Nagai, 1994:309, pl.147.
Lucanus
bruanti LACROIX & BOMANS; Araya, 2001:10-11, figs.8-10.
·
India : Assam
·
Bhutan (British Bootan) : Padong. 1914, col. L. Durel, col. R. Oberthur
(male HOLOTYPUS of Lucanus bruanti LACROIX & BOMANS, 1973, #667A en
Museum National d’Histoire Naturelle, Paris)
|
|
|
Male HOLOTYPUS of Lucanus bruanti LACROIX
& BOMANS, 1973, from Bhutan (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus (Lucanus) cambodiensis DIDIER, 1925.
Lucanus cambodiensis DIDIER, 1925:222 [Cambodia : Lao Kay].
Lucanus cambodiensis DIDIER; Didier, 1926:169
Lucanus cambodiensis DIDIER; Didier & Séguy, 1952:pl.17
Lucanus cambodiensis DIDIER; Benesh, 1960:131
Lucanus cambodiensis DIDIER; Maes, 1992:15.
Lucanus cambodiensis DIDIER; Araya, 2001:10-11, figs.11-12.
·
Cambodia : Lao Kay, IX-1919, col. H. Deyrolle (female TYPE of Lucanus
cambodiensis DIDIER, 1925 in Museum National d’Histoire Naturelle, Paris).
|
|
|
Female TYPE of Lucanus cambodiensis DIDIER,
1925, from Cambodia (adapted from Araya, 2001). |
Bibliography
:
ARAYA K.
(2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Lucanus) cantori ssp. cantori HOPE, 1842 [Khasi Hills].
Lucanus
cantori HOPE, 1842:83 [Khasi Hills].
Lucanus
cantori HOPE; Hope, 1843:363
Lucanus
cantori HOPE; Hope, 1845:73
Lucanus
cantori HOPE; Hope & Westwood, 1845:5
Lucanus cantori HOPE; Burmeister, 1847:527
Lucanus cantori HOPE; Planet, 1899:275
Lucanus cantori HOPE; Planet, 1902:57, pl.13
Lucanus cantori HOPE; Heyne & Taschenberg, 1908:50, pl.6
Lucanus cantori HOPE; Gravely, 1915:411
Lucanus cantori HOPE; Didier, 1949:pl.12
Lucanus cantori HOPE; Arrow, 1950:51, pl.3
Lucanus cantori HOPE; Benesh, 1960:131
Lucanus cantori HOPE; Lacroix, 1967:97-98
Lucanus cantori HOPE; Maes, 1992:15-16
Lucanus cantori HOPE; Mizunuma & Nagai, 1994:213, pl.10.
Lucanus cantori HOPE; Araya, 2001:11, fig.13.
·
India : West Bengal : Darjeeling
·
India : West Bengal : Kurseong
·
India : West Bengal : Gopaldhara
·
India : West Bengal : Rungbong Valley
·
India : West Bengal : Darjeeling : Redong
·
India : “?Himalaya”or “Khasi Hills”
(male TYPE of Lucanus cantori HOPE, 1842, # 356 in Oxford
University Museum of Natural History).
·
Bhutan : Pedong
·
Bhutan : Paro Distr.
·
Bhutan : Gedu.
Adults on
? : Castanopsis, Symplocos.
|
|
|
Male TYPE of Lucanus cantori HOPE, 1842
(adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from
Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus (Lucanus) cantori ssp. colasi LACROIX, 1967.
Lucanus cantori ssp. colasi LACROIX, 1967:97-98 [India : Shillong].
·
India : Meghalaya : Shillong (male TYPE of Lucanus cantori ssp.
colasi LACROIX, 1967).
·
India : Meghalaya : Khasi Hills.
Note : it
seems that the nominal subspecies was describe from Khasi Hills, if it is so, colasi
will be synonym of cantori and then the ssp. of Darjeeling is unnamed.
Lucanus
(Lucanus) cervus ssp. cervus (LINNAEUS, 1758).
Scarabaeus
cornutus DALE, 1693:389, pl.17.
Linnaeus,
1745:69; Rossel, 1746:25, pl.4-5
Scarabaeus
cervus LINNAEUS, 1758:353.
Poda,
1761:21; Sulzer, 1761:3, pl.1; Geoffroy, 1762:61, pl.1; Pontopp., 1763:430; Kurz.,
1765:199; Scopoli, 1763:1; De Bomare, 1764:486; Gronov, 1764:147; Muller,
1764:2; Schaeffer, 1766:pl.9; Linnaeus, 1767:559; Schluga, 1767:pl.1; De Geer,
1774:327, pl.12; Fabricius, 1775:1
Lucanus capreolus FUESSLY, 1775:3
Lucanus dorcas MULLER, 1776:52
Sulzer,
1776:19, pl.2; Bergstrasser, 1778:18, 25, pl.4, 13; Fuessly, 1778:63;
Schaeffer, (1766)-1779:pl.133; Barbut, 1781:15, pl.2; Fabricius, 1781:1;
Fuessly, 1781:2; Schrank, 1781:32; Von Laichart., 1781:1; Von Scheven, 1782:60;
Panzer, 1783:36, pl.29
Lucanus
cervus LINNAEUS; De Asso, 1784
Leske,
1784:464; Bonsd., 1785:221; Fourcroy, 1785:2; Von Moll, 1785:149; Fabricius,
1787:1; Petagna, 1787:1; Von Meidinger, 1787:117; Zschach, 1788:1; De Villers,
1789:41
Lucanus capra OLIVIER, 1789:9-12, pl.1-2
Schrank,
1789:374; Von Razoum., 1789:138; Brahm, 1790:124, 130
Lucanus
armiger HERBST, 1790:287, 289, 299, 301, pl.D-E, pl.33-34.
Lucanus
hircus HERBST, 1790:287, 289, 299, 301, pl.D-E, pl.33-34.
Linnaeus
(ed. Gmelin), 1790:1588; Preyssler, 1790:9; Roesel, 1790:1749, pl.iv; Rossi,
1790:1-2; Scriba, 1790:40; Mayer, 1791:181; Donovan, 1792:31, pl.13; Fabricius,
1792:236-237; Kugelann, 1792:255; Martyn, 1792:pl.5; Brahm, 1793:657;
Fabricius, 1793:236; Schwarz, 1793:84; Panzer, 1795:244; Hoppe, 1796:169, 179;
Hoppe, 1797:179, 200; Panzer, 1798:11;
Olivier
(ed. Illiger), 1800:55-59; Paykull, 1800:45; Walther, 1800:352; Fabricius,
1801:248-249
Lucanus inermis MARSHAM, 1802:46, 48, pl.2 [Loc. ?].
Blumenb.,
1803:321; Latreille, 1804:246, pl.86; Posselt, 1804:15, pl.ii; Duftschm.,
1805:64; Panzer, 1805:110; Bechst. & Scharfenb., 1805:80; Sturm,
1805:pl.1-2; Thunberg, 1806:190-191; Fallen, 1807:12, 22
Lucanus
grandis HAWORTH, 1807:77
Latreille,
1807:135; Gyllenhall, 1808:65-66; Schonherr, 1817:318, 320-321; W.S. Macleay,
1819:114; Blot, 1824:84; Muller, 1825:553-672; Klug, 1829:365, pl.15; Stephens,
1830:166; Davis, 1832:86; Curtis, 1834:490; Audoin, 1835:81; Pictet, 1836:393;
Dejean, 1837:193; Mulsant, 1839:119, pl.2; Newport, 1839:853-994; Ratzeburg,
1839:105; Westwood, 1839:187; Castelnau, 1840:171; Spry & Shuckard,
1840:34, pl.42; Heer, (1838)-1842:495
Lucanus
microcephalus MULSANT, 1842:8, 583, 586, pl.3
La Ferté Senectère en Mulsant, 1842:8;
Burmeister, (1835)-1843:pl.24; Sturm, 1843:347, pl.5; Tulk, 1843:451; Brulle,
1844:271-374; Desmarest, 1845:53
Lucanus
americanus HOPE & WESTWOOD, 1845:9-10 [N. America, error]
Lucanus
lusitanicus HOPE & WESTWOOD, 1845:9-10 [Portugal].
Lucanus
vicinus HOPE & WESTWOOD, 1845:9-10 [Poonah, error].
Motschulsky,
1845:60; Bellier de la Chavignerie, 1846:28; Blanchard, 1846:322
Lucanus
major BURMEISTER, 1847:350, 527
Erichson,
1848:936, 938; Kriechbaun., 1848:208;
Reiche,
1853:71; Perris, 1854:105; Mulsant & Godart, 1855:250; Mulsant &
Godart, 1855:150; Lacordaire, 1856:23; Reiche, 1856:81; Delarouzee, 1859:98;
Delarouzee, 1859:42; Duval, 1859:7, 9-10, pl.1; Gerst., 1859:110; Deyrolle,
1860:22; Kraatz, 1860:68, 265, 273, pl.7; Parry, 1864:71; Abeille de Perrin,
1867:65, 70; Cornelius, 1867:435; Cornelius, 1868:33; Landois, 1868:88-95, pl.vi;
MacLachlan, 1868:ix; Motschulsky, 1870:25-31, 34, pl.2; De Narcillac, 1871:186;
Hochuth, 1871:85; Mulsant & Rey, 1871:8, 14, 16; Schioedte, 1873:342-345,
pl.15; Redtenbacher, 1874:451; Schiodte, 1874:341, pl.xviii-xix; Schlechtendal,
1874:25; Schlechtendal, 1875:158; Kraatz, 1876:72; Rossi, 1876:95;
Schmidt-Goebel, 1876:392; Perris, 1877:101-102, pl.5; Rossi, 1878:227;
Schreiber, 1878:259; Brandt, 1879:1-17; Cotton, 1880:83-84; Reichenau, 1880:55;
Rupertsberger, 1880:142; Hammond, 1881:14-26, pl.2; Reichenau, 1881:172-194;
Kraatz, 1883:27; Leboeuf, 1883:45; Strubing, 1883:159; Cornelius, 1884:33;
Smith, 1885:27; Frivaldsky, 1886:78; Gadeau de Kerville, 1886:176; Fallou,
1889:141; Rousseau, 1889:170; Fowler, 1890:4; Planet, 1890:156-157, 164-165,
215-216
Lucanus
antennatus REY, 1890:164
Reitter,
1891:142-143; Verhoeff, 1893:113-170; Verhoeff, 1893:209-260; Binet,
1894:449-580; Pottier, 1894:27; Rupertsberger, 1894:146; Borgschulze, 1895:131;
Caradja, 1895:77; Eggers, 1895:131; Schaufuss, 1895:111; Selmons, 1895:52;
Planet, 1896:11, 99, 168, 238, 256; Planet, 1897:64, 192, 229, 267; Weise,
1897:389; Bothe, 1899:140; Planet, 1899:31, 35, 42, 46, 55, 58, 62, 64-65, 104,
107, pl.9, 10, 14; Rupertsberger, 1899:235; Stichel, 1899:19
Lucanus
longipennis VON ROTHENBURG, 1900:99
Jarder,
1901:28; Le Senechal, 1901:190; Xambeau, 1901:604; Planet, 1902:52, 100, 111,
114-115, 119, 120, 125; Weber, 1902:335; Biedermann, 1903:395-481
Lucanus
cervus LINNAEUS; Champion, 1903
Lucanus
cervus LINNAEUS; Champion, 1904
Marchand,
1904:37; Rudow, 1904:179, 187, 196; Schuster, 1904:338; Amman, 1905:139;
Scabra, 1905:7-8, pl.; Rey, 1906:59; Barthe, 1909:5-6; Chagrin, 1909:404;
Reitter, 1909:299, pl.68; Chinaglia, 1910:1-8, 25-39; Leoni, 1910:153
Lucanus
capitalis MOLLENKAMP, 1910:242, 575
Schieser,
1910:29; Bedel, 1911:8; Kapzov, 1911:297-337; Krizenecky, 1911:113; Pic,
1911:399; Sasse, 1911:69-104
Lucanus validus MOLLENKAMP, 1912:302 [Borneo, error].
Muzik, 1912:189; Ramme, 1912:37; Roubal,
1912:386; Sirguey, 1912:134; Boileau, 1913:218, 220; Bordas, 1913:358-364;
Fairmaire & Planet, 1913:195, pl.9; Kuhnt, 1913:380; Bordas, 1914:181-183;
Braatz, (1913)-1914:267; Moritz, 1914:138; Stellwag, 1914:359-429; Chinaglia, 1915:7;
Houlbert & Monnot, 1915:4, 6; Labitte, 1916:105; Schaufuss, 1916:1252,
pl.48; Verhoeff, 1916:354-372; Della Beffa, 1921:3; Houlbert (1912)-1921:167;
Dudich, 1923:129; Grutzner, 1924:130; Nagel, 1924:398-399; Henriksen,
1925:134-143; Nagel, 1928:273
Lucanus
ornatus BERNAU, 1929:72
Catherine,
1929:5-6; Vignon, 1929:89-125; Gadeau de Kerville, 1930:63; Nagel, 1930:90;
Goldschmidt, 1931
Lucanus
nigrinus MARCH, 1931.
Marcu,
1931:214; Portevin, 1931:2-3, pl.6; Roubal, 1931:36; Baudrimont, 1932:116;
Franz, 1932:134; Joy, 1932:242; Joy, 1932:pl.69; Kraemer, 1932:321; Kraemer,
1932:86-117; Porta, 1932:371; Didier, 1933:24, pl.1; Hepp, 1933:126-127; Hepp,
1934:84; Golovyanko, 1935:38; Van Emden, 1935:194, 200; Hepp, 1936:173;
Paulian, 1936:751-759; Didier, 1937:xxxviii, xlii; Muller & Schoop,
1937:41-116; Palm, 1941:17; Van Emden, 1941:119; Garnier, 1945:110; Sellier
& Razet, 1947:223-225; Colas, 1949:128-131; Didier, 1949:pl.1, 3;
Leiler,
1950:157; Muller & Schoop, 1950:17-18; Arrow, 1951:pl.8; Condrillier,
1951:20; Brull, 1952; Didier & Séguy, 1952:pl.15, 17, 24; Tressens,
1952:57; Auber, 1953:99-100; Didier & Séguy, 1953:27-32; Tippmann, 1954;
Theodorides, 1955; Britton, 1956:6; Landin, 1956:1; Paulian, 1956:36; Freude,
1957:129-134; Harz, 1957:22-23; Dorn, 1958:62-64
Lucanus cervus LINNAEUS; Español, 1958
Lucanus cervus (LINNAEUS); Horion, 1958
Papperitz, 1958; Weise, 1958; De Ligondes,
1959:52-56; Franz, 1959:74-79; Marie, 1959:125-127; Paulian, 1959:38-39;
Weinreich, 1959:282-283; Benesh, 1960:132, 138-140; Janssens, 1960:95;
Muspratt, 1960:53; Kuhn, 1961; Colas, 1962:118; Demoulin, 1963:pl.8
Lucanus cervus f. scapulodonta WEINREICH, 1963:29.
Burton,
1965:121; Clark, 1966:199
Lucanus
cervus LINNAEUS; Bahuena, 1967
Clark,
1967:24; Lacroix, 1967:98
Lucanus
cervus (LINNAEUS); Freude, 1968:59-64
Lacroix,
1968:234-235; Lacroix, 1969:119-129; Machatschke, 1969; Mathieu, 1969:1061;
Freude, 1971:283
Lucanus
cervus (LINNAEUS); Cofais, 1972:32-33
Lovari,
1972:36
Lucanus
cervus (LINNAEUS); Mal, 1972:161-167
Lacroix
& Bomans, 1973:152; Kersmaekers, 1973:219; Verstraeten, Boosten &
Gaspar, 1973:mapa 713; Jacobs & Renner, 1974; Radisic, 1974; Harde,
1975:16; Athias, 1976:194; Nadolski, 1976; Andersson, 1977:100
Lucanus
cervus (LINNAEUS); Clark, 1977:9
Leech,
1977:69; Perrier, 1977:167; Hempel & Schiemenz, 1978; Lacroix, 1978:253;
Cameron, 1980:31; De Walsche, 1980:201; Krikken, 1980:24
Lucanus
cervus (LINNAEUS); Roer, 1980:248-251
Numhauser,
1981:74; Boosten, 1982:96; Decelle, 1982:203; Español & Belles, 1982:73;
Klausnitzer, 1982:14, 17; Bessonnat, 1984:77-80; Lojtnant, 1984; Fellenberg,
1985:79-80; Stolk, 1985:159; Bartolozzi, 1986:12; Cruysbergh, 1986:103;
Chalmers-Hunt, 1987:184; Creutz, 1987:58-59; Long, 1987:323-324; Marples,
1987:149-150; Newton, 1987:371; Sakaino, 1987:16; Alexay, 1988:24-25;
Andersson, Coulianos, Ehnstrom, Hammarsted, Imby, Janzon, Lindelow &
Walden, 1987:70; Gjurasin, 1988:60-61; Grayson, 1988:224; Rabil, 1988:217; Verdcourt,
1988:36; Wootton, 1988:113; Bartolozzi, 1989:104; Carriere, 1989:19-20; George,
1989:28; Pfaff, 1989:31-32; Baker, 1990:12; Hein, 1990:34; Verdcourt, 1990:196
Lucanus
cervus (LINNAEUS); Zemp, Hartmann, Knecht & Sprecher, 1990.
Bowdrey,
1991:300; Gouranton, Folliot & Thomas, 1991:102-109; Kranjcev, 1991:18-19;
Kuhnel, 1991:209; Mamonov, 1991:157-163; Morris, 1991:106; Newton, 1991:36-38;
Shepard & Biggs, 1991:7-8; Delatour, 1992:75; Alvarez-Lao, 1992:316-323;
Maes, 1992:16-17; Spooner, 1992:329-330; Baraud, 1993:61; Bartolozzi &
Onore, 1993:126
Lucanus
cervus (LINNAEUS); Ochse, 1993:47-52.
Whitehead,
1993:206; Wuest, 1993:132; Hecq, 1994:455; Karner, 1994:71-72; Mizunuma &
Nagai, 1994:212, pl. 8-9
Lucanus
cervus (LINNAEUS); Klausnitzer, 1995.
Viejo & Sanchez Cumplido, 1995:186;
Bartolozzi, 1996:486; Teunissen & Smit, 1996:134-135; Araya, 1996:19;
Köteles & Bokony, 1996:99; Kral & Vitner, 1996:425
Lucanus cervus (LINNAEUS); López-Colón, González Peña & Beltrán
Valen, 1996:16, 18.
Lucanus cervus cervus (LINNAEUS); López-Colón, González Peña & Beltrán
Valen, 1996: 17.
Cecchi & Bartolozzi, 1997:122;
Heynderycx, 1997:392-393, 396; Hoekstra, 1997:93-95.
Lucanus
cervus LINNAEUS; Sprecher-Uebersax & Durrer, 1998:142-166.
Lucanus
cervus LINNAEUS; Sprecher-Uebersax & Durrer, 1998:471-479.
Lucanus
cervus LINNAEUS; Sprecher-Uebersax & Durrer, 2001:2-11.
Lucanus
cervus LINNAEUS; Sprecher-Uebersax & Durrer, 2001:161-182.
Lucanus
cervus (LINNAEUS); Reyes-Castillo, 2002:486.
Lucanus
cervus (LINNAEUS); Ballerio, 2003:68, 73, 82.
Norway
Sweden
Finland
Denmark
Great
Britain : Borders (Eccles--55°37N-2°22W), Northumberland
(Horsley--54°59N-1°09W), Cumbria (Keswick--54°36N-3°07W), W. Yorks
(Burley--53°55N-1°41W, Apperley--53°51N-1°42W), Lancashire
(Thornton--53°53N-3°01W), Clwyd (Prestatyn--53°20N-3°23W), Lincoln
(Lincolnshire--53°13N-0°31W, Boston--52°59N-0°00--Billups), Nottingham
(Nottingham--52°58N-1°09W), Derby (Derbyshire--52°56N-1°28W, Calke,
Calke--Garneys), Shropshire (Wellington--52°40N-2°30W {Ercall wood, Apley wood,
Ollerton wood}, Church Stretton--52°33N-2°49W), Hereford & Worcester
(Bewdley--52°22N-2°20W, Bewdley--52°22N-2°20W--Blatch,
Worcestershire--52°11N-2°14W), Warwick (Warwick--52°18N-1°34W,
Warwick--52°18N-1°34W--Martineau), Cambridge (Cambridgeshire--52°12N-0°08E--Bloomfield),
Buckingham (Buckinghamshire--52°01N-1°00W), Suffolk (Woodbridge--52°06N-1°18E,
Ipswich--52°04N-1°09E, Nayland--51°58N-0°52E, Felixstowe--51°58N-1°20E), Essex
(Halstead--51°57N-0°21E, Manningtree--51°56N-1°04E, Colchester--51°54N-0°54E,
Braintree--51°53N-0°33E, Coggeshall--51°52N-0°41E, Marks Tey--51°52N-0°48E,
Brightlingsea--51°49N-1°01E, Witham--51°48N-0°39E, West Mersea--51°47N-0°55E,
Epping Forest--51°42N-0°06E, Brentwood--51°37N-0°18E, Tilbury--51°27N-0°22E),
Gloucester (Cheltenham--51°54N-205°W, Birdlip--51°49N-2°07W,
Newnham--51°48N-2°28W, Hardwicke--51°48N-2°18W), Hertford
(Hatfield--51°47N-0°14W, Waltham Cross--51°42N-0°11W), Oxford
(Oxford--51°46N-1°15W, Wallingford--51°36N-1°09W, Mapledurham Park), London
(Enfield-- 51°39N-0°05W,
Pinner--51°36N-0°24W, Harrow--51°35N-0°21W = Harrow North,
Ilford--51°34N-0°08E, Romford--51°34N-0°11E, Uxbridge--51°33N-0°29W,
Upminster--51°33N-0°15E, London--51°32N-0°15W, Southall--51°31N-0°23W,
Dagenham--51°31N-0°01E, Woolwich--51°29N-0°03E, Greenwich--51°28N-0°01E,
Eltham--51°27N-0°03E, Hounslow--51°25N-0°21W, Hampton--51°25N-0°21W = Hampton
Court, Kingston--51°25N-0°18W, Streatham--51°25N-0°08W, Bromley--51°25N-0°01E,
Sidcup--51°25N-0°07E, Wimbledon--51°23N-0°12W,
Croydon--51°22N-0°06W, Sutton--51°21N-0°11W, Purley--51°20N-0°07W, Hatton
Garden, Cadogan Square, New Forest), Glamorgan (Swansea--51°37N-3°56W),
Buckingham (Marlow--51°35N-0°46W), Berkshire (Cookham--51°34N-0°42W,
Maidenhead--51°31N-0°43W {Kimbers--Spring Hill}, Slough-- 51°31N-0°36W,
Pangbourne--51°29N-1°07W {Tidmarsh}, Windsor Forest--51°29N-0°37W,
Datchet--51°29N-0°35W, Caversham-- 51°28N-0°59W
= Caversham Warren, Reading--51°27N-0°58W {Tilehurst}), Kent
(Dartford--51°26N-0°14E, Gravesend--51°26N-0°22E, Rochester--51°25N-0°35E, Sheppey
I.--51°25N-0°50E {Sheerness--51°26N-0°46E}), Surrey (Staines--51°26N-0°29W,
Chertsey--51°24N-0°29W, Surbiton--51°24N-0°19W, Weybridge--51°23N-0°35W,
Walton--51°23N-0°35W, Bagshot--51°22N-0°42W = Bagshot Heat,
Epsom--51°20N-0°17W, Woking--51°19N-0°25W, Leatherhead--51°18N-0°20W, Box
Hill--51°16N-0°20W, Merrow--51°15N-0°32W, Godalming--51°12N-0°37W,
Chiddingford--51°07N-0°39W, Merton Park, Kew, Great Leight Rectory), Hampshire
(Farnborough--51°17N-0°46W, Winchester--51°05N-1°19W, Bitterne--50°55N-1°22W, Southampton--50°55N-1°24W,
Havant--50°52N-0°59W, New Forest--50°51N-1°35W, Beaulieu Abbey--50°49N-1°28W,
Gosport--50°48N-1°08W, Lymington--50°46N-1°33W, Southsea--50°46N-1°04W),
Somerset (Castle Cary--51°06N-2°31W), Wilts (Salisbury--51°05N-1°48W), Devon (Instow--51°03N-4°10W,
Merton--50°53N-4°06W = Merton Park), W. Sussex (Midhurst--50°59N-0°45W,
Arundel--50°51N-0°35W, Chichester--50°50N-0°48W, Shoreham-by-See--50°50N-0°17W
= Shoreham, Worthing--50°49N-0°22W, West Tarring), E. Sussex
(Brighton--50°46N-0°08W), Dorset (Lyme Regis--50°43N-2°57W), Cornwall
(Yelverton--50°29N-4°05W, Saltash--50°24N-4°12W, Polperro--50°19N-4°31W,
Truro--50°14N-5°03W), Middlesex (Teddington--Fulwell Vicarage, Heston,
Lampton), Redmarley, Little Tring, Puttenham, Appleslade, Ellingham, Highwood,
Linwood, Calmore, Hamble, Brightwell, Purbrook, Baughurst, Tadley, Mapledurham,
Tidmarsh, Ewelme, Swyncombe Down, Bray, Henley, Eashing, Frimley, Wellington
College, Brookwood, Sunninghill, Stoke Podges, Buryan, Fulwell, Mount Edgcumb,
Bere Alston, Buckland Monachorum, Fingle Gorge, Slapton, Colyton, Parkestone,
West Howe, Mudeford, Bransgore, West Cowes, Alverstoke, Kirby Bedon,
Broxbourne, Chesnut, Wormley, Great Leighs, West Stow, Layer Breton,
Layer-de-La-haye, Peldon, Tollesbury, Stanway, Boxted, Great Horkesley, Mount
Bures, East Mersea, Fingringhoe, Arleigh, Elmstead, East Donyland,
Thorpe-le-Soken, Wivenhoe, Dedham, Lawford, Thorrington Street, Kersey, Great
Bentley, Little Bentley, Bradfield, Brantham, Holbrook, Woolverstone, Little Oakley,
Guilford, Wisley, Oatlands, Shepperton, Northwood, Lancing, Sompting, Bramber,
Henfield, Steyning, Brockham, Ashtead, Bookham, Ranmore, Chessington, Claygate,
East Molesley, Esther, Hanworth, New Malden, Teddington, Fetcham, Heston, Kew,
Lampton, Petersham, Richmond Park, Sheen, Carshalton, Mitcham, Barnes,
Battersea, Cadogan Square, Chelsea, Chiswick, Fulham, Putney, Roehampton Lane,
Wandsworth, Wandsworth Common, Addiscombe, Beckenham, Beddington, Norbury,
Stoneleigh, Blackheat, West Dulwich, Bostall Heat, Brockley, Catford,
Camberwell, Lambeth, Lee, Lewisham, South Norwood, Southwark, Sydenham, Hatton
Garden, Holborn, Chingford, Barcombe, Abbey Wood, Charlton, Chislehurst, Danson
Park, Plumstead Common, Upper Belvedere, Leyton, Loughton, Theydon Bois,
Lullingstone, Darenth Park, Harold Wood, Larkfield, Grays, Stifford, West
Thurrock, Aylesford, Birling, Ditton;
Netherlands
: Drenthe, Overijssel--Twente, Gelderse Vallei, Utrecht (Utrechtse Heuvelrug),
Noord-Brabant, Zuid-Limburg, Eysden, Alpesdroom;
Belgium :
Anvers (Merxem, Postel), Limbourg (Lanaken--50°53N-5°39E,
Vliermaalroot--50°50N-5°23E, Canne), Brabant (Louvain--50°52N-4°42E,
Uccle--50°49N-4°21E, Overijse--50°46N-4°34E, Halle--50°44N-4°14E, Boistfort,
Hoeilart, Bois de la Cambre, Etterbeek, Auderghem, Rouge-Cloître, Watermael,
Boendael, Notre-Dame-au-Bois, Linkebeek, Rhode-Sainte-Genèse, Buysinghen,
Assche), Liège (Wandre--50°40N-5°40E, Liège--50°38N-5°35E,
Jupille--50°38N-5°38E, Chênée--50°37N-5°37E, Chaudfontaine--50°34N-5°40E,
Huy--50°31N-5°15E, Sart--50°31N-5°57E, Spa--50°29N-5°52E,
Aywaille--50°28N-5°40E, Bois d'Aywaille--50°28N-5°40E--Kin, Fléron, Ougrée,
Embourg, Chèvremont, Beaufays, Tilff, Strée, Bois l'Evèque, Val Benoît,
Jemeppe, Cointe, Beyne, Méry), Hainaut (Chimay--50°03N-4°19E--Forges-lez-Chimay,
Harmignies, Vergnies), Namur (Namur--50°28N-4°52E, Hastière--50°14N-4°42E,
Eprave, Foy-Notre-Dame, Waulsort, Marche-les-Dames), Luxembourg
(Saint-Hubert--50°02N-5°23E, Bouillon--49°47N-5°05E, Laroche, Izel, Mardesous);
Luxembourg (Echternach--49°49N-6°26E--Berdorf);
France : Nord (Armentières--50°41N-2°52E),
Calvados (Saint Aubain--49°20N-0°23W, Pont l'Evêque--49°17N-0°11E, Bures),
Aisne (Vise), Marne (Chalons-sur-Marne--48°58N-4°22E), Meuse, Yvelines
(Poissy--48°55N-2°05E, Saint Germain en Laye--48°52N-2°05E, Draveil, Jubisy,
Bois de Boulogne, Auteuil, Longchamps, Achères, Neuilly, Vincennes, Vésinet,
Croissy, Bougival, Marly), Val D'Oise (Montmorency--48°59N-2°22E),
Hauts-De-Seine (Meudon--48°50N-2°15E), Paris (Paris--48°50N-2°20E), Essonne
(Brétigny-sur-Orge--48°37N-2°18E), Finistère (Benodet--47°53N-4°07W), Morbihan
(Grandchamp--47°46N-2°51E), Seine-Et-Marne (Melun--48°33N-2°38E--Mée), Terr. de
Belfort (Delle--47°31N-6°59E), Indre-Et-Loire (Tours--47°25N-0°42E),
Loir-Et-Cher (Romorantin--47°22N-1°44E), Cher (Meillant), Côte d'Or (Petigny),
Allier (Saint Ennemond, Saint Evremond, Tronçais), Saône et Loire (Les
Antoines, Issy l'Evêque), Charentes Maritimes (Montguyon--45°11N-0°12W),
Charente (Féolle, Floirac), Dordogne, Limousin, Loire (Saint
Etienne--45°28N-4°25E), Isère, Haute-Savoie (Chamonix--45°55N-6°51E,
Annecy--45°54N-6°07E), Gironde (Bordeaux--44°50N-0°37W, La Ruscade),
Lot-Et-Garonne (Agen--44°11N-0°40E), Tarn-Et-Garonne (Puylaroque,
Montauban--44°00N-1°12E), Aveyron (Decazeville--44°33N-2°14E, Saint-George de
Luzençon), Ardèche (Casteljau, Saint-Remèze, bois de Païolive), Drôme (Saint
Donat sur l'Herbasse--45°07N-4°59E), Landes (Parentis--44°21N-1°07W, Dax--43°42N-1°02W),
Gers (Escorneboeuf), Pyrénées-Atlantiques (Salies de Bearn--43°29N-0°58W, Saint
Jean de Luz--43°21N-1°26W, Pau--43°18N-0°25W), Hautes Pyrénées
(Lourdes--43°06N-0°02E, Cauterets--42°52N-0°03W, Lespoune), Ariège
(Pamiers--43°06N-1°37E, Foix--42°03N1°36E), Tarn (Grésigne), Aude (Esperanza),
Pyrénées Orientales (La Massane, Ria, Banyuls), Hérault
(Montpellier--43°36N-3°57E, Saint Gély, Saint Guilhem, Minerve), Bouches du
Rhône (Auriol--43°22N-5°38E, Marseille--43°17N-5°24E, Géménos--43°17N-5°37E,
Saint Pons), Vaucluse (Mts. du Luberon--43°50N-5°20E, Oppéde), Alpes de Haute
Provence (Banon--44°02N-5°37E, Riez--43°49N-6°05E), Var (Le Luc--43°24N-6°19E,
La Roquebrussane--43°20N-5°59E, Mts. Sainte Baume--43°19N-5°45E, Forêt de
Sainte-Baume--43°19N-5°45E, Pignans--43°18N-6°14E, Signes--43°17N-5°50E,
Montrieux le Vieux--43°15N-5°58E, Collobrières--43°15N-6°19E,
Belgentier--43°14N-6°01E, Toulon--43°07N-5°55E, Hyères--43°07N-6°04E, Correns,
forêt de Morrières, Evenos, Aiguines, Mt. Touar, Valbelle), Alpes Maritimes
(Saint Cézaire, Gilette), Corse--42°N-9°E, ? (Guyon, Saint Martin
Vesubie);
Spain :
Cantabria : Potes, 43°09 N - 4°37 W
Spain :
Guipúzcoa : Tolosa, 43°09 N - 2°05 W
Spain :
Cataluña
Spain :
Huesca : Torla, 42°38 N - 0°06 W
Spain :
Huesca : Benasque, 42°36 N - 0°31 W
Spain : Huesca : Valle de Gistain, 42°36 N -
0°20 E
Spain : Huesca : Plan, 42°35 N - 0°20 E
Spain :
Huesca : Jaca, 42°34 N - 0°33 W
Spain :
Huesca : Loarre, 42°19 N - 0°37 W
Spain : Huesca : La Peña
Spain : Teruel : Albarracín, 40°30 N - 1°34
W
Germany : Hamburg (Hamburg--53°32N-10°00E),
Nordrhein-Westfalen (Dortmund--51°31N-7°27E), Hessen (Wetzlar--50°33N-8°31E,
Kassel--51°19N-9°28E), Rheinland (Koblenz--50°22N-7°36E), Baden-Wurtemberg
(Pforzheim--48°55N-8°42E), Rostock, Schwerin, Neubrandenburg, Magdeburg,
Potsdam (Velten--52°41N-13°11E), Berlin, Frankfurt, Halle, Cottbus, Erfurt,
Leipzig, Dresden, Suhl, Gera, Chemnitz = Karl-Marx-Stadt, ? (Kauden,
Bentheumer Woud);
Poland;
Czech
Rep. : Vinnéjezero, Pálava Biosphere Reserve;
Slovakia;
Switzerland
: Basel : Kleinhuningen, 47°35 N – 7°35 E, 252 m, 1955 (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Basel : Riehen, 47°35 N – 7°39 E : Schaferstrasse, 281 m, 1988 (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Basel : Bachgraben, 275 m (citated by Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Basel : Baumlihof, 259 m (citated by Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Basel : Lange Erlen, 253 m, 1965 (citated by Sprecher-Uebersax & Durrer,
1998).
Switzerland
: Basel : Schwarz-Park, 261 m, 1992 (citated by Sprecher-Uebersax & Durrer,
1998).
Switzerland
: Basel : Bettingen, 47°34 N – 7°40 E : Chrischona, 470 m (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Basel : Basel, 47°33 N – 7°35 E (citated by Sprecher-Uebersax & Durrer,
1998).
Switzerland
: Bern : Augst, 47°32 N – 7°43 E : Schlossli, 274 m, 1989 (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Bern : Augst, 47°32 N – 7°43 E : Schonbuhl, 289 m, 1965 (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Bern : Munchenstein, 47°31 N – 7°37 E : Hofmatt, 273 m, 1984 (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Bern : Munchenstein, 47°31 N – 7°37 E : Ob. Wannen, 332 m, 1990 (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Bern : Muttenz, 47°31 N – 7°39 E : Schanzli, 260 m, 1991 (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Bern : Muttenz, 47°31 N – 7°39 E : Rutihard, 370 m, 1990 (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Bern : Giebenacht, 47°31 – 7°48, 318 m, 1992 (citated by Sprecher-Uebersax
& Durrer, 1998).
Switzerland
: Bern : Therwil, 47°30 N – 7°33 E : Froloo, 391 m, 1942 (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Bern : Reinach, 47°30 N – 7°35 E : Leiwald, 310 m (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Bern : Arlesheim, 47°30 N – 7°37 E : Spitalholz, 348 m, 1990 (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Bern : Arlesheim, 47°30 N – 7°37 E : Schwimmbad, 300 m (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Bern : Arlesheim, 47°30 N – 7°37 E : Reichenstein, 460 m (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Bern : Lausen, 47°29 N – 7°45 E, 344 m, 1984 (citated by Sprecher-Uebersax
& Durrer, 1998).
Switzerland
: Bern : Blauen, 47°27 N – 7°31 E : Weekendhauser, 570 m (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Bern : Bubendorf, 47°27 N – 7°44 E : Wildenstein, 475 m (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Bern : Zwingen, 47°26 N – 7°32 E : Bahnhof, 339 m, 1995 (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Bern : Himmelried, 47°26 N – 7°35 E: Steffen, 574 m, 1995 (citated by
Sprecher-Uebersax & Durrer, 1998).
Switzerland
: Bern : Windentalhohe, 420 m, 1989 (citated by Sprecher-Uebersax & Durrer,
1998).
Switzerland
: Bern : Grammont, 539 m, 1989 (citated by Sprecher-Uebersax & Durrer,
1998).
Switzerland
: Bern : Schleifenberg, 534 m, 1990 (citated by Sprecher-Uebersax & Durrer,
1998).
Switzerland
: Bern : Uetental, 380 m (citated by Sprecher-Uebersax & Durrer, 1998).
Switzerland : Jura : Lajoux, 47°17 N – 7°08
E
Switzerland
: Vaud : Genève, 46°12N-6°09E
Switzerland
: Bagatz
Osterreich
: Ost-Tirol : Thuringen, 47°12 N - 9°46 E
Osterreich
: Donau : Neuburg Bergheim
Osterreich : Wien, 48°12 N-16°21 E
Hungary :
Mt. Bukk, 48°04N-20°29E
Hungary : Gödölló, 47°37N-18°54E
Hungary : Veszprém, 47°05N-17°57E
Hungary :
Lake Balaton, 46°50N-17°45E : Tihany
Hungary :
Mecsek Mts., 46°10N-17°56E
Hungary :
Zebegewy
Hungary :
Pomaz
Hungary :
Borosjenö
Hungary :
Torokbalint
Romania
Italy : Piemonte (Arona--45°45N-8°32E,
Biella--45°34N-8°01E), Lombardia (Varese--45°49N-8°51E--Montvalle,
Como--45°48N-9°05E), Venezia Giulia (Udine--46°05N-13°12E,
Udine--46°05N-13°12E--Toppo, Vittorio--45°58N-12°17E, Pordenone--45°57N-12°38E,
Treviso--45°39N-12°14E, Trieste--43°40N-13°46E {San Sergio-Petrigna, Valle
Ostra}), Liguria (Genova--44°22N-8°55E), Toscana (Firenze--43°47N-11°16E =
Florence, Siena--43°19N-11°19E, Cortona--43°15N-12°00E, Badia Prataglia),
Emilia Romagna (La Lama), Sardegna--40°10N-9°E, Sicilia--37°35N-14°15E, ?
(Zattaglia);
Slovenija: Mackovci--46°47N-16°10E,
Maribor--46°34N-15°37E, Celje--46°15N-15°14E, Celje--46°15N-15°14E--Ljubno ob
Savinja River, Ljubljana--46°04N-14°29E, Litija--46°03N-14°52E--Hotic,
Grosuplje--45°57N-14°40E--Tabor, Ajdovscina--45°53N-13°53E,
Cerknica--45°48N-14°20E, Postojna--45°47N-14°11E--Javorniki,
Sezana--45°43N-13°50E, Kocevje--45°39N-14°49E, Kozina--45°34N-13°56E,
Bohinj--Ribcev laz, Bohinj--Vogar, ob Suhi-Bohinj, Bohinjska Bistrica, Crna
prst, Gameljne, Gorica, Kamnik, Komenda, Kostanjevica, Kuzma, Lubnik,
Orehovci--Gornja Radgona, Pekel--Zalec, Podgorje--Slavnik, Radenci, Rakek,
Rakov Skocjan, Senozece, Sorica, Sorsko polje, Tacen, Tatre, Trzin, Vaneca,
Vipava, Vodice, Zg. Tuhinj;
Bosnia & Hercegovina : Hercegovina
(Konjice--43°39N-17°56E, Mostar--43°20N-17°48E);
Yugoslavia : ? (Karlovak);
Bulgaria;
Greece :
Karpenision--38°55N-21°46E;
Turkey;
Russia :
to Ural and Caucasus (Terek River, Khosta);
Iran.
Note :
this species is protected by Bern Convention appendix 3; Habitat Direction
appendix 2; Regional Law of Toscana, Italia, appendix A and B (Ballerio, 2003).
Xylophagous
:
Betulaceae
: Alnus, Carpinus.
Fagaceae : Quercus, Fagus.
Hippocastanaceae : Aesculus.
Juglandaceae : Juglans.
Malaceae
: Pyrus, Malus.
Moraceae
: Morus.
Oleaceae
: Fraxinus.
Pinaceae : Picea, Pinus.
Rosaceae : Cerasus, Prunus.
Salicaceae : Salix, Populus.
Tiliaceae : Tilia.
Ulmaceae : Ulmus.
Saproxylophagous :
Aceraceae : Acer.
Fagaceae
: Quercus.
Can breed
in compost (artificial soil).
Natural enemies :
Birds : Corvidae : Pica.
? : Caprimulgus europaeus, Strix aluco.
Acarina : Uropodidae : Polyaspis
patavinus.
? : Isacis cuspidat, Isacis lucani, Megaselia
rufipes, Monocercomonas melolonthae.
Bibliografía :
BAGUENA L. (1967) Scarabaeoidea de la fauna íbero-balear y pirenaica. C.S.F.C.
Madrid, 1955 (publicado en 1967): 1-567.
BALLERIO A. (2003) EntomoLex : la conservazione degli insetti e
la legge. Memorie Soc. entomol. Italiana, 82(1):17-86.
COFAIS M. (1972) Un cas tératologique chez Lucanus cervus.
L’Entomologiste,
28:32-33.
CLARK
J.T. (1977) Aspects of variation in the sag beetle. Systematic Entomology,
2:9-16.
FREUDE
H. (1968)
Uber Mandibel-Anomalitaten bei Lucanus cervus. Nachrichtenblatt der
Bayerischen Entomologen, 17:59-64.
HORION
A. (1958)
Faunistik der mitteleuropaischen Kafer 6: Lamellicornia. 343 pp.
KLAUSNITZER
B. (1995)
Die Hirschkafer. Neue Brehm-Bucherei, 551:109 pp.
LOPEZ-COLON
J.I., GONZALEZ PEÑA C.F. & BELTRAN VALEN J.R. (1996) Familia : Lucanidae. Catalogus de la
Entomofauna Aragonesa, No. 12:15-20.
MAL N. (1972) Sur le polymorphisme antennaire d’une population provençale de
Lucanus cervus. L’Entomologiste, 38:161-166.
OCHSE
M. (1993)
Angaben zum Hirschkafer, Lucanus cervus L., bei Kassel. Hessische
faunistische Briefe, 13:47-52.
REYES-CASTILLO P. (2002) 27. Lucanidae. Pp. 485-492. En LLORENTE
BOUSQUETS J. & MORRONE J.J. Biodiversidad, Taxonomía y Biogeografía de
Artrópodos de México : Hacia una síntesis de su conocimiento. Universidad
Nacional Autónoma de México. Vol. III:692 pp.
ROER
H. (1980)
Zur Verbreitung und Bestandesdichte des Hirschkafers im Rheinland. Acta Musei Rigenaehradecensis, Serie A, Supplement:
248-251.
SPRECHER-UEBERSAX
E. & DURRER H. (1998) Uber das Vorkommen des Hirschkafers (Lucanus
cervus L.) in der Region Basel (Coleoptera). Mitt. Ent. Ges. Basel,
48(4):142-166.
SPRECHER-UEBERSAX
E. & DURRER H. (1998) Untersuchungen zum Stridulationsverhalten der
Hirschkafer-Larven (Lucanus cervus L.) (Coleoptera : Lucanidae).
Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 71:471-479.
SPRECHER-UEBERSAX
E. & DURRER H. (2001) Beobachtungen zur Nahrungswahl des Hirschkafers
(Lucanus cervus L.) (Coleoptera). Mitteilungen der Entomologischen Gesellschaft
Basel, 51(1):2-11.
SPRECHER-UEBERSAX
E. & DURRER H. (2001) Verhaltenstudien beim Hirschkafers mittels
Telemetrie und Videoaufzeichnungen (Coleoptera, Lucanus cervus L.).
Mitteilungen der Naturforschenden Gesellschaften beider Basel, 5:161-182.
ZEMP
M., HARTMANN K., KNECHT D. & SPRECHER-UEBERSAX E. (1990) Arlesheimer
Wald Spitalholz 1990. Unveroffentliches Manuskript zuhanden der
Kober-Schwabe-Stiftung Arlesheim, 59 pp.
Lucanus
(Lucanus) cervus ssp. fabiani MULSANT & GODART, 1855 [Platycerus].
Hexaphyllus
pontbrianti MULSANT 1839 [Lucanus, Platycerus] [France : Lyon : Bois de
Rochecardon].
Lucanus
pentaphyllus REICHE 1853 [Platycerus] [Gallia meridionalia : circa Telonem et
Portum Veneris].
Lucanus heptaphyllus ABEILLE DE PERRIN 1867.
Lucanus
reichei MOTSCHULSKY 1870 [France : Toulouse].
Lucanus capreolus hexaphyllus (PLANET) DIDIER & SEGUY 1953 [Italie] (nomen
nudum).
SE. France : Rhône (Lyon--45°45N-4°51E--bois
de Rochecardon), Ardèche, Drôme (Nyons--44°21N-5°08E), Landes
(Dax--43°42N-1°02W), Pyrénées Orientales (Prades--42°37N-2°25E,
Port-Vendres--42°29N-3°07E, Ria, Vernet-les-Bains, Banyuls-sur-Mer), Hérault
(Montpellier--43°36N-3°57E), Var (La Roquebrussane--43°20N-5°59E, Montrieux le
Vieux--43°15N-5°58E, Toulon--43°07N-5°55E, Hyères--43°07N-6°04E), Alpes
Maritimes, Corse (St. Hyacinthe, Ponte Vecchiu near Ota).
Note :
For Lacroix (1968) fabiani is only a morfa of cervus because
cervus and fabiani are living in the same region.
Lucanus (Lucanus) cervus ssp.
mediadonta LACROIX, 1978.
Georgia :
Tiflis*.
Lucanus (Lucanus)
cervus ssp. tauricus MOTSCHULSKY, 1845.
Lucanus
maxillaris MOTSCHULSKY 1845 [Crimée, Caucase, Tourcménie].
Lucanus
brevicollis MOTSCHULSKY 1870 [SW European Russia : Voronège].
Lucanus
europaeus MOTSCHULSKY 1870 [Kharcov].
Russia :
Voronesk;
Ucrania :
Kharkov, Krym = Crimea*;
Turkmenistan.
Lucanus (Lucanus) cervus ssp. judaicus
PLANET 1902.
Turkey :
Aïn-Tab* at 90 km NE Alep.
Lucanus
(Lucanus) cervus ssp. turcicus STURM, 1843 [Platycerus].
Romania :
Bucaresti--44°25N-26°07E--Ilfov;
Bulgaria
: Balchik--43°24N-28°09E, Arkutino, Lozenec, Skorpilovce;
Greece : Eubée I.--38°35N-23°50E;
Turkey : Istanbul--41°02N-28°56E =
Constantinople, Istambul--41°02N-28°56E--Forest de Belgrad,
Bursa--40°14N-29°01E, Taurus--Adana--36°56N-35°20E, Alyana.
Lucanus (Lucanus) cervus ssp.
akbesianus PLANET, 1896.
Lucanus cervus turcicus ab. syriacus MOTSCHULSKY (nec MOTSCHULSKY) 1897.
Lucanus
cervus ab. motschulskyi BENESH 1960.
Lucanus poujadei PLANET 1897 [Syrie].
Lucanus smyrniacus DIDIER & SEGUY 1953 [Asie mineure].
Lebanon :
Djezin;
Syria :
Akbes* = Sgée, Marah, Smyrne, Alexandrette, Ghazir, Marsch.
Lucanus
(Lucanus) convexus LACROIX, 1982.
Lacroix, 1982:14;
Maes, 1992:17.
India :
Nagaland*.
Lucanus
(Lucanus) cyclommatoides DIDIER, 1928.
Lucanus cyclommatoides DIDIER, 1928:51
Lucanus cyclommatoides DIDIER; Didier & Séguy, 1953:81
Lucanus cyclommatoides DIDIER; Benesh, 1960:140
Lucanus cyclommatoides DIDIER; Maes, 1992:17.
Lucanus cyclommatoides DIDIER; Araya, 2001:12, figs.16-20.
·
Vietnam : Tonkin : Chapa, 13-VII, col. Jeanvoine (male TYPE of Lucanus
cyclommatoides DIDIER, 1928 en Museum National d’Histoire Naturelle,
Paris).
·
Vietnam : Tonkin : Chapa, 26-IX-1918, col. Jeanvoine, ex. coll. Boileau
(female COTYPE of Lucanus cyclommatoides DIDIER, 1928 en Museum National
d’Histoire Naturelle, Paris).
|
|
|
|
Male TYPE of Lucanus cyclommatoides DIDIER,
1928, from Vietnam : Chapa (adapted from Araya, 2001) |
Female COTYPE of Lucanus cyclommatoides DIDIER,
1928, from Vietnam : Chapa (adapted from Araya, 2001) |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Lucanus) delavayi FAIRMAIRE, 1887.
Lucanus
delavayi FAIRMAIRE, 1887:27 [China : Yunnan].
Lucanus
delavayi FAIRMAIRE; Fairmaire, 1887:98
Lucanus
delavayi FAIRMAIRE; Planet, 1900:228
Lucanus
delavayi FAIRMAIRE; Planet, 1902:68
Lucanus
delavayi FAIRMAIRE; Heyne & Taschenberg, 1908:50
Lucanus
delavayi FAIRMAIRE; Didier, 1949:pl.10
Lucanus
delavayi FAIRMAIRE; Benesh, 1960:140
Lucanus
delavayi FAIRMAIRE; Kurosawa, 1978:143
Lucanus
delavayi FAIRMAIRE; Bomans, 1989:9
Lucanus
delavayi FAIRMAIRE; Maes, 1992:17
Lucanus delavayi FAIRMAIRE; Mizunuma & Nagai, 1994:216, pl.16.
Lucanus delavayi FAIRMAIRE; Araya, 2001:13, figs.25-26.
·
China : Yunnan, col. R.P. Delavay (male
PARATYPE of Lucanus delavayi FAIRMAIRE, 1887, # 1666 in Museum
d’Histoire Naturelle, Geneva).
·
China : Yunnan :
Ta-Pin-Tze
·
China : Yunnan :
Yunnan-Sen
·
China : Yunnan :
San-Nen-Kai.
|
|
|
male PARATYPE of Lucanus delavayi FAIRMAIRE,
1887, from China : Yunnan (adapted from Araya, 2001) |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Lucanus) deuveianus BOUCHER, 1998.
Lucanus
deuveianus BOUCHER, 1998:182-184, fig. 2, 4 [China : Guangxi : Daming Shan].
Lucanus
deuveianus BOUCHER; Araya, 2001:14, figs.29-30.
·
China : Guangxi : Daming Shan, 1200 m, VI-1995, col. T. Deuve (male
HOLOTYPUS of Lucanus deuveianus BOUCHER, 1998 in Museum National
d'Histoire Naturelle, Paris).
|
|
|
male HOLOTYPUS of Lucanus deuveianus BOUCHER,
1998, from China : Guangxi : Daming Shan (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus (Lucanus) didieri PLANET, 1927.
Lucanus didieri PLANET, 1927:106 [China : Yunnan : Pe-Yen-Tsin]
Lucanus didieri PLANET; Didier & Séguy, 1952:pl.15
Lucanus didieri PLANET; Benesh, 1960:140
Lucanus didieri PLANET; Lacroix & Bomans, 1973:145
Lucanus didieri PLANET; Maes, 1992:17
Lucanus didieri PLANET; Mizunuma & Nagai, 1994:309, pl.147.
Lucanus didieri PLANET; Araya, 2001:14, figs.31-32.
·
China : Yunnan :
Pe-Yen-Tsin, ex. col. Le Moult (male TYPUS of Lucanus didieri PLANET,
1927 in Museum National d'Histoire Naturelle, Paris).
|
|
|
male TYPUS of Lucanus didieri PLANET,
1927, from China : Yunnan : Pe-Yen-Tsin (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Lucanus) dohertyi BOILEAU, 1911.
Lucanus
laminifer var. minor WATERHOUSE 1890.
Waterhouse,
1890:33; Boileau, 1911:435, pl.34; Arrow, 1950:56; Didier & Séguy, 1952:pl.15; Benesh, 1960:140; Lacroix
& Bomans, 1973:138, 142; Lacroix, 1982:13; Téllez, 1990:39; Maes, 1992:17;
Mizunuma & Nagai, 1994:214, 309, pl.11.
India :
Assam, Meghalaya (Khasi Hills), Nagaland (Naga Hills*).
Lucanus (Lucanus) elaphus FABRICIUS, 1775.
Lucanus cervus var. DEGEER 1774:33.
Lucanus elaphus FABRICIUS, 1775:2 [Virginia].
Lucanus elaphus FABRICIUS; Fabricius, 1781:2.
Lucanus elaphus FABRICIUS; Fabricius, 1887:1.
Lucanus elaphus FABRICIUS; Olivier, 1789:12-13, pl.3.
Lucanus
elaphus FABRICIUS; Linnaeus (ed. Gmelin), 1790:1589.
Lucanus
elaphus FABRICIUS; Fabricius, 1792:237.
Lucanus
elaphus FABRICIUS; Olivier (ed. Illiger), 1800:60-61.
Lucanus
elaphus FABRICIUS; Fabricius, 1801:249.
Lucanus
elaphus FABRICIUS; Olivier (ed. Sturm), 1802:2, pl.2.
Lucanus
elaphus FABRICIUS; Thunberg, 1806:191.
Lucanus
elaphus FABRICIUS; Schonherr, 1817:322.
Lucanus elaphus FABRICIUS; Dejean, 1837:193.
Lucanus elaphus FABRICIUS; Burmeister, 1847:354.
Lucanus elaphus FABRICIUS; Fuchs, 1882:49.
Lucanus elaphus FABRICIUS; Albers, 1890:88.
Lucanus elaphus FABRICIUS; Planet, 1902:92, pl.16.
Lucanus elaphus FABRICIUS; Ulke, 1902.
Lucanus
elaphus FABRICIUS; Wickham, 1903:205-206.
Lucanus
elaphus FABRICIUS; Heyne & Taschenberg, 1908:50, pl.6.
Lucanus
elaphus FABRICIUS; Blatchley, 1910:905.
Lucanus
elaphus ab. carlengi ANGELL 1916:70 [Kentucky : Louisville].
Lucanus
elaphus FABRICIUS; Brimley, 1938.
Lucanus
elaphus FABRICIUS; Loding, 1945.
Lucanus
elaphus FABRICIUS; Didier, 1949:pl.11.
Lucanus
elaphus FABRICIUS; Benesh, 1960:140-141.
Lucanus
elaphus FABRICIUS; Arnett, 1965.
Lucanus
elaphus FABRICIUS; Ritcher, 1966.
Lucanus
elaphus FABRICIUS; Kirk, 1969.
Lucanus
elaphus FABRICIUS; Kirk, 1970.
Lucanus
elaphus FABRICIUS; Arnold & Drew, 1987.
Lucanus elaphus FABRICIUS; McNamara, 1991.
Lucanus elaphus FABRICIUS; Ratcliffe, 1991:267.
Lucanus elaphus FABRICIUS; Maes, 1992:17.
Lucanus elaphus FABRICIUS; Anonymous, 1994.
Lucanus elaphus FABRICIUS; Mizunuma & Nagai, 1994:212, pl. 7.
Lucanus
elaphus FABRICIUS; Staines, 1994:17-18.
Lucanus
elaphus FABRICIUS; Poole & Gentili, 1996:297.
Lucanus
elaphus FABRICIUS; Staines, 2000:26-30.
Lucanus
elaphus FABRICIUS; Staines, 2001:397-404.
Canada
:
Ontario (citated by Benesh, 1960).
USA :
Minnesota (National Museum of
Natural History, Washington).
Michigan (Ohio State University).
Nebraska : Richardson Co. : Stella,
6-VI-1999 (University of Nebraska).
Iowa : Lee Co. : Keokuk (Museum of
Comparative Zoology, Cambridge).
Kansas (Field Museum of Natural
History, Museum of Comparative Zoology, Cambridge, National Museum of Natural
History, Washington).
Kansas : Douglas Co. (University of
Kansas).
Kansas : Labette Co. : Oswego
(National Museum of Natural History, Washington).
Kansas : Leavenworth Co. :
Leavenworth, 23-VII-1943 (California Academy of Science).
Kansas : Leavenworth Co. :
Leavenworth, 30-VII-1947 (University of Kansas).
Kansas : Wilson Co. : New Albany
(Natural History Museum, London).
Kansas : Wyandotte Co. (Field Museum
of Natural History).
Missouri (Field Museum of Natural
History).
Missouri : Boone Co. : Columbia,
VI-1903 (Field Museum of Natural History).
Missouri : Boone Co. : Columbia,
21-VI-1971 (University of Missouri).
Missouri : Boone Co. : Columbia,
14-VII-1959 (University of Missouri).
Missouri : Boone Co. : Columbia,
2-VIII-1968 (University of Missouri).
Missouri : Boone Co. : Columbia,
4-IX-1965 (University of Missouri).
Missouri : Butler Co. : 20-VI-1897
(National Museum of Natural History, Washington).
Missouri : Poplar Bluff, VII-1952
(University of Missouri).
Missouri : Girardeau Co. : Cape
Girardeau, 25-VII-1956 (National Museum of Natural History, Washington).
Missouri : Jasper Co. (IMRM).
Missouri : Jefferson Co. : Barnhart,
30-VII-1960 (University of Missouri).
Missouri : Jefferson Co. : Barnhart,
3-VIII-1935 (University of Missouri).
Missouri : New Madrid Co. :
Morehouse, 6-V-1915 (Museum of Comparative Zoology, Cambridge).
Missouri : Pike Co. : VII-1934
(University of Missouri).
Missouri : Louisiana, 6-VI-1923
(University of Missouri).
Missouri : Platte Co. : 29-VI-1992
(University of Missouri).
Missouri : Pulaski Co. : 17-VII-1956
(Florida State Collection of Arthropods).
Missouri : St. Charles Co. :
O'Fallen, 4-VI-1904 (University of Missouri).
Missouri : St. Louis Co. : Creve
Coeur, 1-VII-1908 (University of Missouri).
Missouri : Meramec Hinds, 3-VI-1923
(National Museum of Natural History, Washington).
Missouri : St. Louis, 20-VII-1897
(National Museum of Natural History, Washington).
Missouri : St. Louis, 20-VI-1904
(Field Museum of Natural History).
Missouri : St. Louis, 23-VI-1907
(University of Michigan).
Missouri : St. Louis, 28-VI-1972
(University of Missouri).
Missouri : St. Louis (California
Academy of Science, Carnegie Museum of Natural History).
Missouri : Stoddard Co. : Popular
Bluff Forestry Camp, 23-VI-1954, taken at light (Florida State Collection of
Arthropods).
Missouri : Stone Co. : 20-VIII-1915
(University of Missouri).
Missouri : Taney Co. : Forsyth,
14-VII-1929 (Field Museum of Natural History).
Missouri : Texas Co. : Mountain
Grove, 1916 (National Museum of Natural History, Washington).
Missouri : Washington Co. :
Irondale, 29-VII-1961 (University of Missouri).
Missouri : Wayne Co. :
Williamsville, 20-III-1961 (University of Missouri).
Missouri : Wright Co. : Cedar Gap,
5-VI-1911 (Field Museum of Natural History).
Illinois (University of Minnesota).
Illinois : Southern Illinois
(National Museum of Natural History, Washington).
Illinois : Atlas, VI-1931
(University of Missouri).
Illinois : Adams Co. : Quincy
(Illinois Natural History Survey).
Illinois : Alexander Co. : Cairo
(National Museum of Natural History, Washington).
Illinois : Villa Ridge, IX-1892
(Illinois Natural History Survey).
Illinois : Calhoun Co. : Batchtown,
8-VI-1899 (Illinois Natural History Survey).
Illinois : Clay Co. : Flora
(National Museum of Natural History, Washington).
Illinois : Edwards Co. : 15-VI-1944
(Illinois Natural History Survey)
Illinois : Hamilton Co. : Dahlgren,
4-VI-1885 (Illinois Natural History Survey).
Illinois : Knox Co. : Galesburg
(Museum of Comparative Zoology, Cambridge)
Illinois : Platt Co. : Allerton
Estate, 26-VI-1959 (Illinois Natural History Survey).
Illinois : Randolph Co. : 24-V-1896
(University of Missouri).
Illinois : Richland Co. : 3-VI-1926
(Illinois Natural History Survey).
Illinois : Wabash Valley, V-1878
(Museum of Comparative Zoology, Cambridge)
Illinois : St. Clair Co., 21-VI-1902
(University of Missouri)
Illinois : St. Clair Co.,
1-VIII-1902 (University of Missouri)
Illinois : Manssa, 30-V-1921
(Illinois Natural History Survey).
Illinois : Union Co. : Anna,
17-VII-1883 (Illinois Natural History Survey)
Illinois : Pine Hills Recreation
Area, 10-VII-1968, blacklight trap (Florida State Collection of Arthropods).
Illinois : Washington Co. : Dubois,
1909 (Illinois Natural History Survey).
Indiana (National Museum of Natural
History, Washington).
Indiana : Clay Co. : Saline City,
6-VI-1889 (National Museum of Natural History, Washington).
Indiana : Crawford Co. (citated by
Blatchley, 1910).
Indiana : Daviess Co. : Bicknell,
4-VI-1937 (University of Arkansas).
Indiana : Daviess Co. :
Bicknell, 6-VI-1938 (University of
Arkansas).
Indiana : Dubois Co. : 13-VIII-1970
(Texas A & M University).
Indiana : Elkhart Co. : Elkhart
(California Academy of Science).
Indiana : Knox Co., 27-V-1944
(University of Arkansas).
Indiana : Knox Co., 10-VI-1943
(University of Arkansas).
Indiana : Monroe Co. : Bloomington,
VI-1928 (California Academy of Science)
Indiana : Monroe Co. : Bloomington,
13-VI-1931 (California Academy of Science)
Indiana : Orange Co. : Orleans,
3-VII-1935 (University of Arkansas).
Indiana
: Vigo Co. (citated by Blatchley, 1910).
Indiana : Wabash Co. : Mt. Vernon,
23-V-1964 (Florida State Collection of Arthropods).
Indiana :
Corydon (in coll. Jay Timberlake, USA).
Ohio (National Museum of Natural
History, Washington).
Ohio : Bowlin, VII-1937 (Ohio State
University).
Ohio : Erie Co. : Sandusky (Cedar
Point), 23-VII-1904 (California Academy of Science).
Ohio : Hamilton Co. : Cincinnati
(National Museum of Natural History, Washington).
Kentucky (Illinois Natural History
Survey).
Kentucky : Cumberland Gap (Museum of
Comparative Zoology, Cambridge).
Kentucky : Oragus (Museum of
Comparative Zoology, Cambridge).
Kentucky : Bell Co. : Balkan,
VI-1915 (National Museum of Natural History, Washington).
Kentucky : Henderson Co. : Henderson,
25-V-1921 (Field Museum of Natural History).
Kentucky : Jefferson Co. :
Louisville, 38°15N-85°45W, 9-VII-1938 (University of Kentucky)
Kentucky : Letcher Co. : Whitesburg,
VIII-1915 (National Museum of Natural History, Washington).
Kentucky : Meade Co. : Otter Creek
Park, 19-VI-1970 (National Museum of Natural History, Washington).
Kentucky : Todd Co. : Elkton, 17
June 1961 (University of Kentucky).
Kentucky : Warren Co. : Bowling
Green, VII-1892 (University of Kentucky).
Pennsylvania (Field Museum of
Natural History).
Pennsylvania : Delaware Co. :
Lansdowne, 10-VI-1935 (California Academy of Science).
Pennsylvania : Greene Co. :
Mapletown, 4-I-1961 (National Museum of Natural History, Washington).
Pennsylvania : Montgomery Co.
(Museum of Comparative Zoology, Cambridge).
West Virginia : Logan Co. : Fort
Branch, 27-V-1911 (National Museum of Natural History, Washington).
West Virginia : Marion Co. :
Fairmont, 1926 (Illinois Natural History Survey).
West Virginia : Mingo Co. : Laurel Creek
Public Hunting and Fishing Area, 14-VI-1979 (West Virginia University).
Virginia (TYPUS de Lucanus elaphus FABRICIUS)
Virginia : Buckingham Co. :
27-V-1911 (National Museum of Natural History, Washington).
Virginia : Lunenburg Co. : Victoria
(National Museum of Natural History, Washington).
Virginia : Montgomery Co. :
Blacksburg, 5-V-1956 (Virginia Polytechnic Institute).
Virginia : Nelson Co., 9-VII-1914
(National Museum of Natural History, Washington).
Virginia : Nelson Co., 5-VIII-1927
(National Museum of Natural History, Washington).
Virginia : Pittsylvania Co. :
Chatham, 26-VII-1939 (Virginia Polytechnic Institute).
Virginia : Danville (Field Museum of
Natural History).
Virginia : Prince Edward Co. :
Farmville, 24-V-1923 (University of Georgia).
Virginia : Prince Edward Co. :
Farmville, 24-VI-1924 (University of Georgia).
Virginia : Prince George Co. :
Hopewell, 23-VI-1930 (National Museum of Natural History, Washington).
Virginia : Petersburg Fort Lee,
23-VI-1962 (National Museum of Natural History, Washington)
Virginia : 10 miles south of
Petersburg, 4-VI-1970 (National Museum of Natural History, Washington).
Virginia : Richmond Co. : Warsaw,
16-VI-1938 (National Museum of Natural History, Washington).
Virginia : Sussex Co. : Jarratt,
18-VI (Cornell University).
District of Columbia (citated by
Uike, 1902)
Maryland : Baltimore (Natural
History Museum, London).
Maryland : St. Mary's Co. : Great
Mills, 18-VIII-1947, taken at light (Florida State Collection of Arthropods)
Maryland : St. Mary Co. : Lexington Park, 4-VII-1992,
taken at light (Florida State Collection of Arthropods)
Maryland : Patuxent River Naval Air
Station, 7-VIII-1975 (Philip J. KEAN, USA).
Maryland : Worcester Co. : Pocomoke
City, taken in blacklight trap (Florida State Collection of Arthropods).
Maryland : Worcester Co. : Pocomoke
City, 1896 (University of Maryland).
Delaware : Sussex Co. : Milton,
6-VI-1940 (University of Delaware).
Oklahoma : Choctow Co. : Boswell,
3-VI-1925 (California Academy of Science).
Oklahoma : Craig Co. : Blue Jacket,
30-VI-1931 (Field Museum of Natural History).
Oklahoma : Creek Co. : Dumright,
13-VI-1925 (California Academy of Science).
Oklahoma : Latimer Co., VII-1982
(Florida State Collection of Arthropods).
Oklahoma : Latimer Co., VII-1987
(Florida State Collection of Arthropods).
Oklahoma : Red Oak, VII-1981
(Florida State Collection of Arthropods).
Oklahoma : 5 miles west of Red Oak,
16-VII-1977 (Florida State Collection of Arthropods).
Oklahoma : 5 miles west of Red Oak,
VII-1979 (Florida State Collection of Arthropods).
Oklahoma : Mays Co., 12-VII-1980
(Texas A & M University).
Oklahoma : McCurtain Co., 8-VI-1934
(Field Museum of Natural History).
Oklahoma : McCurtain Co.,
11-VII-1932 (Field Museum of Natural History).
Oklahoma : 6.5 miles northeast of
Idabel, 28/29-VIII-1937 (University of Michigan).
Oklahoma : Mclntosh Co. (citated in
Arnold and Drew, 1987).
Oklahoma : Okmulgee Co. : Haskell,
VII-1905 (National Museum of Natural History, Washington).
Oklahoma : Osage Co. : Bartlesville
(National Museum of Natural History, Washington).
Oklahoma : Ottawa Co. : Commerce,
VIII-1934 (University of Kansas)
Oklahoma : Wyandotte, 15-VI-1931
(University of Kentucky).
Oklahoma : Payne Co. : Still-water,
18-VII-1928 (California Academy of Science).
Arkansas : Arkansas Co. : 8-VI-1970,
blacklight (University of Arkansas).
Arkansas : Ashley Co. : Hamburg
(University of Arkansas).
Arkansas : Benton Co. : Siloam
Springs, 18-VI-1907 (National Museum of Natural History, Washington).
Arkansas : Columbia Co. : Magnolia,
1931 (University of Kansas).
Arkansas : Devil's Den State Park,
20-VI-1975 (University of Arkansas).
Arkansas : Fayetteville (University
of Arkansas).
Arkansas : Garland Co. : Hot Springs
(National Museum of Natural History, Washington).
Arkansas : Hempstead Co. : Hope,
16-VI-1935 (Museum of Comparative Zoology, Cambridge).
Arkansas : Hempstead Co. : Hope,
19-VI-1931 (Cornell University).
Arkansas : Hempstead Co. : Hope,
VII-1926 (Museum of Comparative Zoology, Cambridge).
Arkansas : Hempstead Co. : Hope,
31-VII-1930 (California Academy of Science).
Arkansas : Independence Co. :
Newark, 23-VI-1886 (National Museum of Natural History, Washington).
Arkansas : Merkel, 26-II-1909
(National Museum of Natural History, Washington).
Arkansas : Johnson Co., 5-VI-1975
(University of Arkansas).
Arkansas : Johnson Co., 23-VI-1977
(University of Arkansas).
Arkansas : Lafayette Co.,
2/3-VI-1965 (University of Arkansas).
Arkansas : Lawrence Co. : no further
data (California Academy of Science).
Arkansas : Imboden (California
Academy of Science).
Arkansas : Imboden, 25-V-1929
(California Academy of Science)
Arkansas : Imboden, 13-VI-1931
(California Academy of Science)
Arkansas : Imboden, 15-VI-1925
(California Academy of Science)
Arkansas : Imboden, VII-1926
(California Academy of Science, Field Museum of Natural History)
Arkansas : Imboden, 13-VII-1925
(Field Museum of Natural History).
Arkansas : Logan Co. : Magazine
Mountain : Cameron Bluff Campground, 28-VI-1990 (American Museum of Natural
History, New York).
Arkansas : Logan Co. : Magazine
Mountain : Cameron Bluff Campground, 18-VII-1990 (American Museum of Natural
History, New York).
Arkansas : Mississippi Co. :
6-VI-1971 (University of Arkansas).
Arkansas : Montgomery Co. : Upper Lake
Ouachita (National Museum of Natural History, Washington).
Arkansas : Montgomery Co. : Upper
Lake Ouachita, 6-VIII-1961 (National Museum of Natural History, Washington).
Arkansas : Phillips Co. : Helena,
16-VI-1920 (Mississippi State University).
Arkansas : Pulaski Co. : Little
Rock, 26-VI-1983 (Florida State Collection of Arthropods).
Arkansas
: Pulaski Co. : Maumalle.
Arkansas : Savoy, 20-VII-1988 (University of Arkansas)
Arkansas : Sebastian Co. : Fort
Smith, 13-VII-1983, light trap (Florida State Collection of Arthropods).
Arkansas : Sebastian Co. : Fort
Smith, 1-VII-July 1984 (Florida State Collection of Arthropods).
Arkansas : Sevier Co., VIII-1972
(University of Arkansas).
Arkansas : St. Francis Co. : Forrest
City, VI-1965 (National Museum of Natural History, Washington).
Arkansas : Washington Co., 1938
(University of Arkansas).
Arkansas : Washington Co., 3-VI-1977
(University of Arkansas).
Arkansas : Washington Co.,
20-VI-1964 (University of Arkansas).
Arkansas : Washington Co.,
30-VII-1971 (University of Arkansas).
Arkansas : Washington Co., 8-X-1976
(University of Arkansas).
Arkansas : Washington Co., 4-XII
(University of Arkansas).
Tennessee (Natural History Museum,
London, California Academy of Science).
Tennessee : Bedford Co. (National Museum
of Natural History, Washington).
Tennessee : Benton Co. (Carnegie
Museum of Natural History, Florida State Collection of Arthropods, National
Museum of Natural History, Washington).
Tennessee : Can-oil Co. (Florida
State Collection of Arthropods, National Museum of Natural History,
Washington).
Tennessee : Coffee Co. (University
of Tennessee).
Tennessee : Davidson Co. (National
Museum of Natural History, Washington).
Tennessee : Dickson Co. (Florida
State Collection of Arthropods).
Tennessee : Gibson Co. (University
of Tennessee).
Tennessee : Hamilton Co. (Florida
State Collection of Arthropods, National Museum of Natural History,
Washington).
Tennessee : Hardeman Co. (CMNM).
Tennessee : Knox Co. (University of
Tennessee, Field Museum of Natural History).
Tennessee : Marion Co. (Carnegie
Museum of Natural History).
Tennessee : Morgan Co. (California
Academy of Science, Carnegie Museum of Natural History, Field Museum of Natural
History, Florida State Collection of Arthropods, Museum of Comparative Zoology,
Cambridge, Manchester Museum, University of Kentucky, National Museum of
Natural History, Washington).
Tennessee : Obion Co. (Illinois
Natural History Survey).
Tennessee : Roane. Co. (Carnegie
Museum of Natural History, Museum of Comparative Zoology, Cambridge).
Tennessee : Rutherford Co.
(University of Michigan).
Tennessee : Sevier Co. (Texas A
& M University).
Tennessee : Shelby Co. (Carnegie
Museum of Natural History).
North Carolina (California Academy
of Science, Field Museum of Natural History, Museum of Comparative Zoology,
Cambridge, Florida State Collection of Arthropods, National Museum of Natural
History, Washington).
North Carolina : Bertie Co. (North
Carolina State University).
North Carolina : Cabarrus Co.
(California Academy of Science, Florida State Collection of Arthropods, North
Carolina State University).
North Carolina : Camden Co. (North
Carolina State University).
North Carolina : Carteret Co.
(Florida State Collection of Arthropods).
North Carolina : Catawba Co. (North
Carolina State University).
North Carolina : Chatham Co. (North
Carolina State University).
North Carolina : Cleveland Co.
(North Carolina State University).
North Carolina : Craven Co. (Florida
State Collection of Arthropods).
North Carolina : Davidson Co. (North
Carolina State University).
North Carolina : Duplin Co. (North
Carolina State University).
North Carolina : Durham Co. (North
Carolina State University).
North Carolina : Edgecomb Co. (North
Carolina State University).
North Carolina : Franklin Co. (North
Carolina State University).
North Carolina :
Gaston Co (Museum of Comparative Zoology, Cambridge, North Carolina State
University).
North Carolina : Graham Co.
(California Academy of Science).
North Carolina : Granville Co.
(North Carolina State University, University of Kansas, University of
Nebraska].
North Carolina : Halifax Co.
(National Museum of Natural History, Washington).
North Carolina : Henderson Co.
(North Carolina State University).
North Carolina : Iredell Co. (North
Carolina State University).
North Carolina : Jackson Co. (Ohio
State University).
North Carolina : Johnston Co. (North
Carolina State University, Florida State Collection of Arthropods).
North Carolina : Macon Co. (North
Carolina State University).
North Carolina : Mecklenburg Co.
(Natural History Museum, London, North Carolina State University).
North Carolina : Moore Co. (North
Carolina State University, Ohio State University).
North Carolina : Nash Co. (National
Museum of Natural History, Washington).
North Carolina : Person Co. (North
Carolina State University, University of Minnesota).
North Carolina : Pitt Co.
(University of Kansas).
North Carolina : Randolph Co.
(Cornell University).
North Carolina : Scotland Co.
(National Museum of Natural History, Washington).
North
Carolina : Shelby Co., 35°15N-81°35W : State Forest
North Carolina : Stanly Co. (Texas A
& M University).
North Carolina : Swain Co. (Cornell
University).
North
Carolina : Union Co. : Waxhaw, 34°55N-80°45W
North Carolina : Union Co. (Carnegie
Museum of Natural History, North Carolina State University).
North Carolina : Wake Co. (Field
Museum of Natural History, Florida State Collection of Arthropods, North
Carolina State University, University of Michigan).
North Carolina : Washington Co.
(Florida State Collection of Arthropods).
North Carolina : Wayne Co. (North
Carolina State University).
South Carolina : Tarplay Gap,
4-VII-1956 (Field Museum of Natural History).
South Carolina : Anderson Co. :
Pendleton, 3-VI-1927 (Manchester Museum).
South Carolina : Bamburg Co. :
IX-1927 (Clemson University).
South Carolina : Calhoun Co. :
Congaree Swamp (Carnegie Museum of Natural History).
South Carolina : Fail-field Co. : Parr
(citated in Kirk, 1970).
South Carolina : Fail-field Co. :
Winnsboro (citated in Kirk, 1970).
South Carolina : Florence Co. :
Florence (citated in Kirk, 1969).
South Carolina : Greenville Co. :
Greer, 3-VII-1931 (Clemson University).
South Carolina : Lancaster Co. :
Richburg, 10-VI-1939 (National Museum of Natural History, Washington).
South Carolina : Laurens Co. :
Enoree (Carnegie Museum of Natural History).
South Carolina : Lexington Co. :
Lexington, 1-VI-1982 (Clemson University).
South Carolina : McCormick Co. :
Willington (citated in Kirk, 1970).
South Carolina : Oconee Co. : CCC
Camp, VII-1943 (National Museum of Natural History, Washington).
South Carolina : Oconee State Park,
18-III-1974, reared- emerged 6-IX-1974 (Texas A & M University).
South Carolina : Pickens Co. :
Clemson, 6-VI-1975 (Texas A & M University).
South Carolina : Pickens Co. :
Clemson, 15-VI-1975 (Texas A & M University).
South Carolina : Pickens Co. :
Clemson, 10-VII-1975 (Texas A & M University).
South Carolina : Pickens Co. :
Clemson, 15-VI-1931 (National Museum of Natural History, Washington).
South Carolina : Pickens Co. :
Clemson, 28-VI-1974 (Florida State Collection of Arthropods).
South Carolina : Richland Co. :
Columbia, 1914 (National Museum of Natural History, Washington).
South Carolina : Richland Co. :
Columbia, 6-V-1916 (National Museum of Natural History, Washington).
South Carolina : Spartanburg Co. :
Spartanburg, 13-VI-1929 (Cornell University).
South Carolina : Sumter Co. :
Wedgefield (citated by Kirk, 1969).
South Carolina : York Co. :
27-VII-1972 (University of Kansas).
Louisiana (National Museum of
Natural History, Washington).
Louisiana : Bossier Parish :
Barksdale Air Force Base, 4/16-VI-1996, all taken at blacklight trap
(Mississippi State University).
Louisiana : Caddo Parish:
Shreveport, 16-VI-1990, on pine tree (Louisiana State Arthropod Collection).
Louisiana : East Baton Rouge Parish
: Baton Rouge, 20-VI-1972 (Louisiana State Arthropod Collection).
Louisiana : East Feliciana Parish: Clinton,
9-VI-1971, on basket oak (Louisiana State Arthropod Collection).
Louisiana : East Feliciana Parish:
Clinton, 29-IV / 2-V-1975 (Louisiana State Arthropod Collection).
Louisiana : Natchitoches Parish :
Kisatchie National Forest, 2 miles east Red Bluff Campground, 17/18-VI-1988
(Louisiana State Arthropod Collection).
Louisiana : Tangipahoa Parish :
Loranger, 20-VIII-1931 (Louisiana State Arthropod Collection).
Louisiana : Webster Parish : Lake
Bistineau State Park, 21-V-1996 (Texas A & M University).
Louisiana : West Feliciana Parish :
Tunica Hills WMA, 2-VI-1995 (Louisiana State Arthropod Collection).
Mississippi (Carnegie Museum of
Natural History, Museum of Comparative Zoology, Cambridge)
Mississippi : Amite Co. (Mississippi
State University).
Mississippi : Bolivar Co.
(University of Mississippi).
Mississippi : Calhoun Co. (National
Museum of Natural History, Washington).
Mississippi : Can-oil Co.
(Mississippi State University).
Mississippi : Chickasaw Co.
(Mississippi State University).
Mississippi : Clay Co. (University
of Michigan).
Mississippi : George Co. (Field
Museum of Natural History).
Mississippi : Grenada Co.
(Mississippi State University).
Mississippi : Jasper Co.
(Mississippi State University).
Mississippi : Kemper Co. (National
Museum of Natural History, Washington).
Mississippi : Lafayette Co.
(Mississippi State University, University of Mississippi, National Museum of
Natural History, Washington).
Mississippi : Lee Co. (Mississippi
State University).
Mississippi : Lincoln Co. (Mississippi
State University).
Mississippi : Lowndes Co.
(Mississippi State University).
Mississippi : Marion Co.
(Mississippi State University).
Mississippi : Oktibbeha Co.
(Mississippi State University, California Academy of Science).
Mississippi : Panola Co. (Mississippi
State University).
Mississippi : Perry Co. (Carnegie
Museum of Natural History).
Mississippi : Pontotoc Co.
(Mississippi State University, Ohio State University).
Mississippi : Rankin Co.
(Mississippi State University, Ohio State University).
Mississippi : Stone Co. (Mississippi
State University).
Mississippi : Tippah Co. (University
of Minnesota).
Mississippi : Union Co. (University
of Mississippi).
Mississippi : Warren Co.
(Mississippi State University).
Mississippi : Washington Co.
(Florida State Collection of Arthropods).
Mississippi : Wayne Co. (Louisiana
State Arthropod Collection).
Alabama : no further data (Museum of
Comparative Zoology, Cambridge).
Alabama : Covington Co. (citated by
Loding, 1945).
Alabama : DeKalb Co., 28-VI-1951
(Illinois Natural History Survey).
Alabama : Lawrence Co. (citated by
Loding, 1945).
Alabama : Lee Co. : Auburn,
1-VI-1924 (Field Museum of Natural History)
Alabama : Lee Co. : Auburn,
20-VI-1925 (California Academy of Science).
Alabama : Lee Co. : Auburn, 2-VII-1927
(California Academy of Science).
Alabama : Madison Co. : Huntsville,
VII-1963 (Florida State Collection of Arthropods).
Alabama : Monte Sano State Park
(Florida State Collection of Arthropods).
Alabama : Monte Sano State Park,
1915 (National Museum of Natural History, Washington).
Alabama : Monte Sano State Park,
VI-1935 (California Academy of Science).
Alabama : Monte Sano State Park,
fall-1976 (National Museum of Natural History, Washington).
Alabama : Monte Sano State Park,
fall-1977 (National Museum of Natural History, Washington).
Alabama : Marshall Co. :
Albertville, VII-1919 (Natural History Museum, London).
Alabama : Mobile Co. : Mobile,
VII-1908 (University of Missouri).
Alabama : Monroe Co. : Haines Island
Park, 26/31-V-1995, black light trap (Mississippi State University).
Georgia : Baldwin Co. :
Milledeeville 23 May 1938 (University of Georgia).
Georgia : Chattahoochee Co. : Fort
Benning, VII-1952 (University of Georgia).
Georgia : Clarke Co. : Athens, various
dates (University of Georgia).
Georgia : Clarke Co. : Athens,
X-1934 (National Museum of Natural History, Washington).
Georgia : Dade Co. : Trenton,
21-VI-1937 (University of Georgia).
Georgia : Fulton Co. : 6-VII-1960
(University of Georgia).
Georgia : Gordon Co : 15-VII-1975
(University of Georgia).
Georgia : Habersham Co. : 5 miles
north of Batesville, 21-VI-1987 (Florida State Collection of Arthropods).
Georgia : Morgan Co. : Madison,
5-VI-1939 (University of Georgia).
Georgia : Rutledge, 14-VII-1940
(University of Georgia).
Georgia : Rutledge, 3-VIII-1952
(University of Georgia).
Georgia : Newton Co. : Covmgton,
30-VI-1949 (University of Georgia).
Georgia : Spalding Co. : Griffin
Experiment Station, 8-VIII-1996,
taken on walk at trail garden (University
of Georgia).
Georgia : Union Co. : Rocky Top
Mountain, VI-1960 (University of Georgia).
Florida (Museum of Comparative
Zoology, Cambridge).
Florida : Gadsden Co. :
Chattahoochee, 31-V-1953, light trap (University of Michigan).
Florida : Hamilton Co. : near
Jasper, 15-VI-1977 (Florida State Collection of Arthropods).
Florida : Liberty Co. : Torreya
State Park, 11-VII-1981, taken in river bottom forest (Florida State Collection
of Arthropods).
Florida : Liberty Co. : Torreya
State Park, 17-VII-1982, taken at uv light (Florida State Collection of
Arthropods).
Texas (Museum of Comparative
Zoology, Cambridge, National Museum of Natural History, Washington).
Texas : Bowie Co. : Texarkana,
V-1962 (Texas A & M University).
Texas : Caddo Co. : Caddo Lake State
Park, 20-V-1998 (Texas A & M University).
Texas : Hunt Co. : Quinlan,
25-VI-1940 (Texas A & M University).
Texas : Marion Co. : Lake O'the
Pines, 12-VI-1979 (Texas A & M University).
Texas : Sabine Co. : 9 miles east of
Hemphill, 14-VI-1990, taken in "beech bottom" (Texas A & M
University).
Texas : SFA Forest Camp, Milam,
VI-1965 (Texas A & M University).
Texas : Travis Co. : Austin
(California Academy of Science).
Bibliography
:
Anonymous (1994) Rare, threatened and endangered animals of.Maryland. Maryland
Department of Natrual Resources. 14 pp.
ARNETT R. H. (1965) The beetles of the United States. Tlie American Entomological
Institute. Ann Arbor 1112 pp.
ARNOLD D.C. & DREW W.A. (1987) The stag beetles of Oklahoma (Coleoptera: Lucanidae). Proceedings
of the Oklahoma Academy of Sciences 67:27-29.
BENESH B. (1960) Lucanidae. Coleopteroruni Catalogus Supplementa. W. Junk. s'Gravenhage. pars 8:1-178.
BLATCHLEY W.S. (1910) An illustrated descriptive catalogue of the Coleoptera or beetles
(exclusive of the Rhynchophora) known to occur in Indiana. The Nature
Publishing Company. Indianapolis. 1386 pp.
BRIMLEY C.S. (1938) The insects of North Carolina. North Carolina Department of
griculture, Division of Entomology. Raleigh, North Carolina. 560 pp.
HAWES C.J. (1998) The stag beetle, Lucanus cervus L. (Coleoptera: Lucanidae)
in Suffolk—a first report. Transactions of the Suffolk Naturalists Society
34:35-49.
KIRK V.M. (1969) A list of beetles of South Carolina. Part I—Northern Coastal
Plain. South Carolina Agricultural Experiment Station Technical Bulletin
1033:1-124.
KIRK V.M. (1970) A list of the beetles of South Carolina. Part 2—mountain,
piedmont, and southern coastal plain. South Carolina Agricultural Experiment
Station Technical Bulletin 1038:1-117.
LODING H.P. (1945) Catalogue of the beetles of Alabama. Geological Survey of Alabama
Monograph 11:1-172.
McNAMARA J. (1991) Family Lucanidae-stag beetles [ Pp. 143-145]. In: Checklist of
beetles of Canada and Alaska (Y. Bousquet, editor). Agriculture Canada,
Research Branch Publication 1861/E.
MIZUNUMA T. & NAGAI S. (1994) The lucanid beetles of the world. Mushi-sha. Tokyo. 337 pp.
RITCHER P.O. (1966) White grubs and their allies. Oregon State University Press,
Corvallis.
STAINES C.L. (1994) A "Monstrous" Lucanus elephus (Coleoptera:
Lucanidae) from Maryland. Maryland Naturalist 38(3-4): 17-18.
STAINES C.L. (2000). A survey for Lucanus elaphus (Fabricius) (Coleoptera: Lucanidae)
in southern Maryland. Maryland Naturalist 43(3-4):26-30.
STAINES
C.L. (2001) Distribution of Lucanus elaphus Linnaeus (Coleoptera : Lucanidae)
in North America. The Coleopterists Bulletin, 55(4):397-404.
ULKE H. (1902).
A list of the beetles of the District of Columbia. Proceedings of the United
States National Museum 25(1275): 1-57.
WICKHAM H.F. (1903)
Gynandromorphism in Lucanus elaphus. Canadian Entomologist, 35:205-206.
Lucanus
(Lucanus) fairmairei PLANET, 1897.
Planet,
1897:265; Planet, 1902:80; Nagel, 1929:117; Arrow, 1950:54, pl.4; Benesh,
1960:141; Kurosawa, 1978:143; Maes, 1992:17; Mizunuma & Nagai, 1994:217,
pl.17.
Myanmar =
Burma : Chin State (N. Chin Hills);
China :
Xizang = Thibet* (Se-Pin-Lou-Chan, Ya-Tcheou), Yunnan;
Thailand
: Chiang Mai (Nan).
Lucanus
(Lucanus) feglini LACROIX & BOMANS, 1973.
Lacroix
& Bomans, 1973:148; Maes, 1992:17.
Afghanistan
: Ghazni*.
Lucanus
(Lucanus) formosanus PLANET, 1899.
Planet,
1899:36; Planet, 1902:42; Mollenkamp, 1913:19; Miwa, 1931:321; Benesh,
1960:141; Lacroix, 1972:54; De Lisle, 1973:137-138; Kurosawa, 1978:143;
Sakaino, 1981:23; Klausnitzer, 1982:45; Hashimoto, 1990:23-26, pl.4; Imanishi,
1990:15; Maes, 1992:17; Ma, 1993:280; Sakaino, Yu & Chu, 1994:122; Mizunuma
& Nagai, 1994:215, pl.13; Kawano, 1997:454.
Taiwan =
Formose* : Taoyuan Distr. (Mt. Rara Shan), Hualien Distr. (Pilu, Hungye near
Juisui), Nantou Distr. (Jiyuetan, Puli = Hori), Kaoshiung Distr. (Liukuei),
Taitung Distr. (Chihpen wenchuan), ? (Meishi, Suisharyo).
Note : Ma
(1993) cites this species from China (Hunan--Wuling Mts.).
Lucanus (Lucanus) formosus DIDIER, 1925.
Lucanus
formosus (BOILEAU in litt.) DIDIER, 1925:220 [Laos : Xieng Khouang].
Lucanus formosus DIDIER; Didier, 1926:153
Lucanus formosus DIDIER; Didier & Séguy, 1953:81
Lucanus formosus DIDIER; Benesh, 1960:141
Lucanus formosus DIDIER; Kurosawa, 1978:143
Lucanus formosus DIDIER; Bomans, 1989:4
Lucanus formosus DIDIER; Maes, 1992:17.
Lucanus formosus DIDIER; Araya, 2001:15, figs.33-37.
·
China : Yunnan : Pe-Yen-Tsing (female COTYPUS of Lucanus formosus DIDIER,
1925 in Museum National d’Histoire Naturelle, Paris).
·
China : Fujian : Kuatun
·
Laos : Xieng Khouang, 15-V-1919 (male TYPUS of Lucanus formosus DIDIER,
1925 in Museum National d’Histoire Naturelle, Paris).
|
|
|
|
male TYPUS of Lucanus formosus DIDIER,
1925, from Laos : Xieng Khouang (adapted from Araya, 2001) |
female COTYPUS of Lucanus formosus DIDIER,
1925, from China : Yunnan : Pe-Yen-Tsing (adapted from Araya, 2001) |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Lucanus) fortunei SAUNDERS, 1854.
Saunders,
1854:46, pl.3; Planet, 1902:85, pl.15; Heyne & Taschenberg, 1908:50; Nagel,
1926:117
Arrow,
1943:134; Didier & Séguy, 1952:pl.18; Benesh, 1960:141; Kurosawa, 1978:143;
Bomans, 1989:4, 9; Maes, 1992:17; Mizunuma & Nagai, 1994:216, pl.16.
China* :
Jiangsu, Shanghai (Shanghai), Jiangxi, Zhejiang = Tche-Kiang (Hang-Tcheou, Mt.
Siming Shan), Fujian = Fokien (Yun-Ling-Shan, San-Chiang, Sanming, Mt. Taiyun
Shan, Kuantun), Guangdong, ? (Tche-Ping--Simeon-Ten).
Lucanus
(Lucanus) fossilis WICKHAM, 1913.
Wickham,
1913; Benesh, 1960:153; Maes, 1992:17; Mizunuma & Nagai, 1994:318.
Fossil :
Miocene of Colorado.
Lucanus
(Lucanus) fryi BOILEAU, 1911.
Lucanus
fryi BOILEAU, 1911:431, pl.34 [Burma : Ruby Mines]
Lucanus
fryi BOILEAU; Arrow, 1950:48, pl.4-5
Lucanus fryi BOILEAU; Didier & Séguy, 1952:pl.17
Lucanus fryi BOILEAU; Benesh, 1960:141
Lucanus fryi BOILEAU; Lacroix, 1971:561
Lucanus fryi BOILEAU; Bomans, 1991:146
Lucanus fryi BOILEAU; Maes, 1992:17
Lucanus fryi BOILEAU; Kawano, 1994:9
Lucanus fryi BOILEAU; Mizunuma & Nagai, 1994:213, pl.11
Lucanus fryi BOILEAU; Kawano, 1997:454.
Lucanus fryi BOILEAU; Araya, 15, figs.38-39.
·
Myanmar = Burma : Ruby Mines, col. Doherty, ex coll. Fry 1905-100 (male
HOLOTYPE of Lucanus fryi BOILEAU, 1911 in Natural History Museum,
London) (male PARATYPE of Lucanus fryi BOILEAU, 1911 in Museum National
d’Histoire Naturelle, Paris).
·
Myanmar : Kambaiti
·
China : Yunnan : Pe-Yen-Tsing
·
Thailand : Chiang Mai : Fang
·
Thailand : Chiang Mai : Fang : Mt. Doi Pahompok
|
|
|
male PARATYPE of Lucanus fryi BOILEAU,
1911, from Myanmar = Burma : Ruby Mines (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Lucanus) gamunus SAWADA & WATANABE, 1960 [gammunus, gammus].
Sawada
& Watanabe, 1960:99; Nomura, 1960:39; Kurosawa, 1978:141-153; Ishida,
1987:5; Kojima, 1987:43; Isida & Fujioka, 1988:3; Araya, 1989:6; Téllez,
1990:39; Nakata, 1991:22-25; Maes, 1992:17; Mizunuma & Nagai, 1994:217,
pl.17; Araya, 1996:20.
Japan :
Tokyo Met. [Izu Is. (Kozujima I.,
Mikurajima I. {Sato})].
Lucanus (Lucanus) gennestieri LACROIX, 1971.
Lucanus (Lucanus) gennestieri LACROIX, 1971:565.
Maes, 1992:17; Mizunuma & Nagai,
1994:213.
Lucanus gennestieri LACROIX; Zilioli, 2002:213.
China : Szechuan (Lou-Tse-Kiang*), Yunnan
(Tse-Kou*).
Bibliography
:
ZILIOLI
M. (2002)
Lucanus victorius n. sp., a new outstanding stag beetle from Sichuan,
China (Coleoptera, Lucanidae). Atti Soc. It. Sci. nat. Museo civ. Stor. Nat. Milano, 143(II):209-213.
Lucanus (Lucanus) hermani DE LISLE, 1973.
Lucanus hermani DE LISLE, 1973:137 [China : Fukien : Sha-Bang].
Lucanus hermani DE LISLE; Kurosawa, 1978:143
Lucanus hermani DE LISLE; Sakaino, 1981:23
Lucanus hermani DE LISLE; Bomans, 1989:4
Lucanus hermani DE LISLE; Maes, 1992:18
Lucanus hermani DE LISLE; Mizunuma & Nagai, 1994:215, pl.14.
Lucanus hermani DE LISLE; Araya, 2001:16-17, figs.45-49.
·
China : Sichuan
·
China : Fujian = Fukien
: Yen-Ping, 19-VIII-1920 (male PARATYPUS, female PARATYPUS de Lucanus
hermani DE LISLE, 1973 in Museum d’Histoire Naturelle, Geneva).
·
China : Fujian = Fukien
: Sha-Bang (male HOLOTYPUS de Lucanus hermani DE LISLE, 1973).
·
China : Fujian : Mt.
Taiyun Shan
·
China : Guangdong
·
China : Hainan I. : Mt.
Wuzhi Shan
|
|
|
|
male
PARATYPUS de Lucanus hermani DE LISLE, 1973, from China : Fujian =
Fukien : Yen-Ping (adapted from Araya, 2001). |
female
PARATYPUS de Lucanus hermani DE LISLE, 1973, from China : Fujian =
Fukien : Yen-Ping (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Lucanus) ibericus MOTSCHULSKY, 1845.
Motschulsky,
1845:60; Burmeister, 1847:349; Duval, 1859:11; Kraatz, 1860:268-271, pl.7;
Deyrolle, 1864:312; Motschulsky, 1870:37-41, 50, pl.2; Reitter, 1891:144;
Planet, 1896:237; Planet, 1897:82, 100, 106, 171, pl.15; Planet, 1899:60,
74-75, 78, 85-88, pl.14; Semenow, 1901:105; Mollenkamp, 1906:32; Barthe,
1909:5; Planet, 1909:173; Boileau, 1913:221, pl.9; Schaufuss, 1916:1253;
Schatzmayr, 1923:8; Didier, 1949:pl.9; Didier & Séguy, 1952:pl.8, 15;
Didier & Séguy, 1953:29; Benesh, 1960:141-142; Lacroix & Bomans,
1973:152; Lacroix, 1978:253; Klausnitzer, 1982:8; Maes, 1992:18; Mizunuma &
Nagai, 1994:212, pl.7.
Lucanus
(Lucanus) ibericus ssp. ibericus MOTSCHULSKY, 1845 [Platycerus].
Lucanus
curtulus MOTSCHULSKY 1845 [Caucase].
Lucanus
barbarossa var. minor BURMEISTER 1847.
Lucanus
tetraodon DUVAL (nec THUNBERG) 1859.
Lucanus
orientalis KRAATZ 1860.
Lucanus
intermedius MOTSCHULSKY 1870 [Armenie : Akhalzik].
Lucanus
piger MOTSCHULSKY 1870 [Georgie : Tiflis].
Lucanus syriacus MOTSCHULSKY 1870 [Liban].
Lucanus tenebrosus MOTSCHULSKY 1870 [Lenkoran].
Lucanus caspica SEMENOW 1901.
Lucanus
ravasinii SCHATZMAYR 1923 [Tomorica].
Albania :
Tomorica;
Greece;
Turkey;
Lebanon;
Syria :
Smyrna, Lenkoran;
Georgia :
Tiflis;
Armenia :
Akhalzik.
Lucanus
(Lucanus) ibericus ssp. macrophyllus (REICHE) KRAATZ, 1860 [Platycerus].
Lucanus
laticornis DEYROLLE 1864 [lorticornis] [Syria : Ararat].
Lucanus
subvelutinus MOTSCHULSKY 1870 [Turkmenie, Astrabad--Perse, Caucase].
Pseudolucanus
busignyi PLANET 1909.
Turkmenistan;
Syria :
Mt. Ararat;
Iran = Perse : Astrabad, Caramania,
Siaret--Kopet-Dagh.
Lucanus
(Lucanus) jeanvoinei DIDIER, 1927.
Lucanus
jeanvoinei DIDIER, 1927:203 [Vietnam : Tonkin].
Lucanus jeanvoinei DIDIER; Didier & Séguy, 1952:pl.17
Lucanus jeanvoinei DIDIER; Benesh, 1960:143
Lucanus jeanvoinei DIDIER; Maes, 1992:18.
Lucanus jeanvoinei DIDIER; Araya, 2001:17, figs.50-54.
·
Vietnam : Tonkin (male HOLOTYPE of Lucanus jeanvoinei DIDIER,
1927 in Museum National d’Histoire Naturelle, Paris).
·
Vietnam : Tonkin : Chapa (male PARATYPE of Lucanus jeanvoinei DIDIER,
1927 in Museum National d’Histoire Naturelle, Paris).
|
|
|
|
male HOLOTYPE of Lucanus jeanvoinei DIDIER,
1927, from Vietnam : Tonkin (adapted from Araya, 2001). |
male PARATYPE of Lucanus jeanvoinei DIDIER,
1927, from Vietnam : Tonkin : Chapa (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Lucanus) kanoi KUROSAWA, 1966.
Kurosawa,
1966:339-342; Kurosawa, 1978:144; Hashimoto, 1990:23, 26, pl.4; Imanishi,
1990:15-17; Maes, 1992:18; Sakaino, Yu & Chu, 1994:122; Sakaino,
1994:18-19; Mizunuma & Nagai, 1994:214, pl.12; Sakaino, 1995:3-7; Boucher,
1998:180.
Lucanus
(Lucanus) kanoi ssp. piceus KUROSAWA, 1966.
Taiwan : Taoyuan Distr. (Mt. Rara San),
Taipei Distr. (Mt. Lalashan*), Ilan Distr. (Kuhsha*, Nan-shan).
Lucanus (Lucanus) kanoi ssp. kanoi KUROSAWA, 1966.
Taiwan : Huanlien Distr. (Pilu), Nantou
Distr. (Sungkang--Tattaka*,
Sunkang, Mt. Hehuanmou, Wushe).
Lucanus
(Lucanus) klapperichi BOMANS, 1989.
Bomans,
1989:4, 9-12; Mizunuma & Nagai, 1994:216.
China :
Fujian (Kuatun*).
Lucanus
(Lucanus) koyamai AKIYAMA & HIRASAWA, 1990.
Lucanus
(Lucanus) koyamai AKIYAMA & HIRASAWA in HIRASAWA &
AKIYAMA, 1990:53.
Bomans,
1991:146; Kawano, 1997:454; Kawano, 1997:25.
Lucanus
koyamai AKIYAMA & HIRASAWA; Mizunuma & Nagai, 1994:7-8, fig.
4a-c, 217, pl.17.
Thailand
: Chiang Mai (Fang, Mt. Doi Pahompok near Fang).
Bibliography.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Lucanus
(Lucanus) kraatzi NAGEL, 1926.
Lucanus
(Lucanus) kraatzi ssp. kraatzi NAGEL, 1926.
Lucanus
(Lucanus) kraatzi NAGEL, 1926:116 [Sse-Tsong].
Didier, 1927:202; Didier & Séguy, 1952:pl.17;
Benesh, 1960:143; Maes, 1992:18; Mizunuma & Nagai, 1994:216, pl.16.
Lucanus kraatzi kraatzi NAGEL; Ikeda, 1997:14-16, figs. 5-6.
China :
E. Yunnan
: Sse-Tsong, 2000 m (TYPUS of Lucanus kraatzi NAGEL).
Yunnan :
Li Kiang.
Lucanus
(Lucanus) kraatzi ssp. giangae IKEDA, 1997.
Lucanus
kraatzi giangae IKEDA, 1997:14-16, figs. a-c, 1-4 [Mt. Pia Oac].
Vietnam
:
N.
Vietnam : Cao Bang Prov. : Mt. Pia Oac (male HOLOTYPUS of Lucanus kraatzi
giangae IKEDA, 2 male PARATYPUS in coll. Ikeda, Japan) (1 male PARATYPUS in
coll. Megumu Sugawara, Japan) (4 male PARATYPUS in coll. Fujita, Japan) (1 male
PARATYPUS in coll. Suzumura, Japan) (12 male PARATYPUS in coll. Katsura, Japan)
(1 male PARATYPUS in coll. Kunio Kume, Japan).
Bibliography
:
IKEDA H.
(1997) A new subspecies of Lucanus kraatzi Nagel (Coleoptera, Lucanidae)
from Northern Vietnam. Gekkan-Mushi, 316:14-16.
Lucanus
(Lucanus) kurosawai SAKAINO, 1995.
Sakaino, 1995:3-7; Araya, 1996:22; Araya,
1996:16.
Taiwan :
Nantou (Sungkang*, Jih-yue-tan*, Mt. Hewang-shan* = Habon, Sung-chuan-kang*),
Taichung (Mt. An-ma Shan*, Wu-ling*).
Lucanus
(Lucanus) laetus ARROW, 1943 (new name for oberthuri).
= Lucanus oberthuri PLANET (nec
PLANET 1896) 1897.
Planet, 1897:171; Planet, 1902; Arrow,
1943:134; Didier, 1949:pl.18; Benesh, 1960:143; Kurosawa, 1978:145-146; Maes,
1992:18; Mizunuma & Nagai, 1994:217, pl.17; Kawano, 1997:25.
China : Xizang = Thibet
(Siao-Lou-Lou-Chan*), Sichuan (Leshan), Yunnan.
Lucanus
(Lucanus) laminifer WATERHOUSE, 1890.
Waterhouse,
1890:33; Waterhouse, 1890:pl.186; Planet, 1899:202; Planet, 1902:53; Heyne
& Taschenberg, 1908:50; Boileau, 1913:221; Pouillaude, 1913:332; Gravely,
1915:411; Didier, 1925:219; Didier, 1926:137; Didier, 1949:pl.17; Arrow,
1950:44, pl.3, 5; Didier & Séguy, 1953:87; Benesh, 1960:143, 147; Lacroix,
1972:54; Kurosawa, 1978:143; Sakaino, 1981:23-24; Klausnitzer, 1982:29; Téllez,
1990:39; Bomans, 1991:146; Maes, 1992:18, 20; Mizunuma & Nagai, 1994:216,
pl.15; Kawano, 1997:454.
Lucanus
(Lucanus) laminifer ssp. laminifer WATERHOUSE, 1890.
India :
Sikkim, West Bengal (Darjeeling), Assam*, Meghalaya (Khasi Hills), Nagaland
(Naga Hills, Kohima), Manipur;
Myanmar =
Burma : ? (Ruby Mines);
China :
Yunnan (Jinghong);
Thailand
: Chiang Rai (Wiangpapao), Chiang Mai (Mt. Doi Pui), Meo--N. Thailand.
Lucanus (Lucanus) laminifer ssp.
vitalisi POUILLAUDE, 1913.
Laos;
Cambodge;
Vietnam :
Tonkin* (Chapa, Lao-Kay, Ha-lang, Lao Kay--Hoang Lien So'n, Sapa--Hoang Lien
So'n).
Lucanus
(Lucanus) ludivinae BOUCHER, 1998.
Lucanus
ludivinae BOUCHER, 1998:178-180, figs. 1, 3 [China : NW Yunnan : E. Gaoligong Shan
: Un Shan : District de Baoshan].
Lucanus
ludivinae BOUCHER; Araya, 2001:17-18, figs.59-60.
·
China : NW Yunnan : E. Gaoligong Shan : Un Shan : District de Baoshan,
2500 m, VII-1998, col. J.M. Bousquet (male HOLOTYPUS of Lucanus ludivinae BOUCHER,
1998 in Museum National d'Histoire Naturelle de Paris).
|
|
|
male HOLOTYPUS of Lucanus ludivinae BOUCHER,
1998, from China : NW Yunnan : E. Gaoligong Shan : Un Shan : District de
Baoshan (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Lucanus) lunifer ssp. lunifer HOPE, 1831 [Himalayas].
Lucanus
lunifer HOPE, 1831 [Himalayas].
Lucanus
lunifer HOPE; Westwood en Royle, 1839:55, pl.9
Lucanus
rugifrons HOPE& WESTWOOD, 1845:4, 9 [Himalayas].
Lucanus lama OLIVIER; Burmeister, 1847:353
Lucanus
lunifer HOPE; Kollar & Redtenbacher, 1848:531, pl.24
Lucanus
lunifer HOPE; Reiche, 1853:71
Lucanus
lunifer HOPE; Parry, 1864:72
Lucanus
lunifer HOPE; Planet, 1898:107
Lucanus
lunifer HOPE; Planet, 1902:12, pl.3
Lucanus
lunifer HOPE; Heyne & Taschenberg, 1908:50, pl.6
Lucanus
lunifer HOPE; Boileau, 1913:218
Lucanus
lunifer HOPE; Stebbing, 1914:70
Lucanus
lunifer HOPE; Gravely, 1915:412
Lucanus
lunifer HOPE; Beeson, 1919:488
Lucanus
lunifer HOPE; Didier, 1949:pl.15
Lucanus
lunifer HOPE; Arrow, 1950:45, pl.3, 5
Lucanus
lunifer HOPE; Didier & Séguy, 1953:82
Lucanus
lunifer HOPE; Benesh, 1960:143
Lucanus
lunifer HOPE; Endrodi, 1968:50
Lucanus
lunifer HOPE; Kurosawa, 1970:160, pl.1
Lucanus
lunifer HOPE; Lacroix, 1971:555, 557
Lucanus
lunifer HOPE; Nakane, 1972:41, 45
Lucanus
lunifer HOPE; Lacroix, 1981:61, 64
Lucanus
lunifer HOPE; Klausnitzer, 1982:42
Lucanus
lunifer HOPE; Lacroix, 1982:16
Lucanus
lunifer HOPE; Sakaino, 1987:16
Lucanus
lunifer HOPE; Arif, Shah & Joshi, 1990:36
Lucanus
lunifer HOPE; Maes, 1992:17, 18
Lucanus
lunifer HOPE; Bartolozzi & Sforzi, 1994:112, 114
Lucanus
lunifer HOPE; Mizunuma & Nagai, 1994:213, pl.10.
Lucanus
lunifer HOPE; Araya, 2001:18, figs. 61-62.
Lucanus
rugifrons HOPE & WESTWOOD; Araya, 2001:19-20, fig.72.
Lucanus
lunifer HOPE; Zilioli, 2002:213.
·
China : Xizang = Thibet
·
China : Yunnan : Djo
Kou La, 31°25 N - 102°58 E
·
India : Jammu & Kashmir
·
India : Himachal Pradesh : Palampur
·
India : Himachal Pradesh : Chamba Distr., 6 km of Tissa
·
India : Punjab : Dalhousie
·
India : Uttar Pradesh = United Provinces : Dehra Dun
·
India : Uttar Pradesh = United Provinces : Mussoorie
·
India : Uttar Pradesh = United Provinces : Naini Tal
·
India : Uttar Pradesh = United Provinces : Simla
·
India : Uttar Pradesh = United Provinces : Josimath
·
India : Sikkim
·
India : West Bengal : Darjeeling
·
India : West Bengal : Kurseong
·
India : West Bengal : Gopaldhara
·
India : West Bengal : Rungbong Valley
·
Himalaya, col. Royle (male TYPE of Lucanus lunifer HOPE, 1831, #
357 in Oxford University Museum of Natural History).
·
Himalaya (female TYPE of Lucanus rugifrons HOPE & WESTWOOD,
1845, #358 in Oxford University Museum of Natural History) [Araya, 2001 wrote
as locality “?Tonkin”].
·
Nepal : Pokhara
·
Nepal : Nimarch
·
Nepal : Sete
·
Nepal : Godavari
·
Nepal : Godavari : Mt.
Pulchok
·
Nepal : Mulkharka :
Kathmandu Valley : Bhaktapur Distr.
·
Nepal : Jiri : Dolaka
Distr.
·
Bhutan : Maria Basti.
Saproxylophagous : Fagaceae : Quercus,
Castanopsis.
Adults on
Poaceae : Zea.
|
|
|
|
|
male TYPE of Lucanus lunifer HOPE,
1831, from Himalaya (adapted from Araya, 2001). |
Female TYPE of Lucanus rugifrons HOPE
& WESTWOOD, 1845, from Himalaya (adapted from Araya, 2001). |
|
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
ZILIOLI
M. (2002)
Lucanus victorius n. sp., a new outstanding stag beetle from Sichuan,
China (Coleoptera, Lucanidae). Atti Soc. It. Sci. nat. Museo civ. Stor. Nat. Milano, 143(II):209-213.
Lucanus (Lucanus) lunifer ssp.
franciscae LACROIX, 1971.
Lucanus
lunifer ssp. franciscae LACROIX, 1971:555, 557 [India : Shillong].
·
India : Assam
·
India : Meghalaya : Khasi Hills
·
India : Meghalaya : Shillong (TYPUS of Lucanus lunifer ssp.
franciscae LACROIX, 1971).
·
Myanmar = Burma.
Lucanus
(Lucanus) maculifemoratus MOTSCHULSKY, 1861.
Motschulsky,
1860:16; Motschulsky, 1861:9; Vollenhoven, 1861:103; Parry, 1862:108; Parry,
1864:9-10, pl.4; Vollenhoven, 1865:147; Motschulsky, 1866:172; Parry, 1870:56;
Parry, 1873:335; Lewis, 1883:333; Heyden, 1884:275; Heyden, 1887:250; Reitter,
1891:144; Jakowleff, 1896:171; Planet, 1897:205; Planet, 1898:19, 251; Planet,
1899:275-276; Planet, 1900:47; Planet, 1902:24, 30, 34-35, 39, 80, 129, pl.5-7,
8, 10; Matsumura, 1906:72, pl.45; Heyne & Taschenberg, 1908:50; Barthe,
1909:5; Mollenkamp, 1912:302; Inukai, 1924:83; Didier, 1926:144; Nagel,
1930:90; Miwa, 1931:318-319; Miwa, 1936:2; Didier, 1949:pl.14; Didier &
Séguy, 1952:223; Didier & Séguy, 1952:pl.21-23; Didier & Séguy,
1953:32, 85; Nakane, 1955:85; Inahara & Baba, 1959:8; Benesh, 1960:141,
143-144, 147; Nomura, 1960:39; Kurosawa, 1964:444; Kurosawa, 1966:341; Nomura,
1966:68; Lacroix, 1968:180; Abe, Kudoh, Kondoh & Saitoh, 1969:180;
Kurosawa, 1970:294; Lacroix, 1972:54; Nakane, 1972:421; Virkki &
Reyes-Castillo, 1972:50; Kurosawa, 1976:189; Kurosawa, 1978:143-144, 148-149;
Nakane, 1979:1; Numhauser, 1981:74; Sakaino, 1981:23; Klausnitzer, 1982:17;
Kikuta, 1984:2; Ishida, 1987:5-9; Nakane, 1987:135; Isida & Fujioka,
1988:3; Araya, 1989:8; Hashimoto, 1990:23-26, pl.4; Imanishi, 1990:15; Araya
& Obuchi, 1992:31; Abe, Kudoh, Ichikawa & Saitoh, 1992:220; Araya,
Obuchi & Ochi, 1992:31; Li, 1992:67-68; Maes, 1992:18-19; Suzuki,
1992:20-23; Ma, 1993:280; Adachi, 1994:18-19; Sakaino, Yu & Chu, 1994:122;
Sakaino, 1994:18-19; Mizunuma & Nagai, 1994:214-215, 215-216, pl.11, 12-13,
14; Tsukawaki, 1995:12-16; Araya, 1996:20; Araya, 1996:16; Kawano, 1997:454.
Lucanus
(Lucanus) maculifemoratus ssp. boileaui PLANET, 1897.
China :
Shaanxi, Sichuan (Mt. Emei Shan), Hubei (Shennongjia, Wuling Mts.), Xizang =
Thibet (Siao-Lou*, Mou-Pin, Se-Pin-Lou-Chan, Ya Tcheou), Yunnan.
Bibliography.
ARAYA
K. (2000)
An account of a visit to European Museums. 2. The lucanid specimens in the van
Roon and Siebold collections deposited in the Rijksmuseum van Natuurlijke
Historie, Leiden. Gekkan-Mushi, 350:4-16.
Lucanus
(Lucanus) maculifemoratus ssp. jilinensis LI, 1992.
China :
Jilin.
Lucanus
(Lucanus) maculifemoratus ssp. dybowskyi PARRY, 1873 [dybouski]
[Thibet, error].
Russia :
Siberia (Amur, Vladivostok, Dauria, Askold I.);
China :
Beijing (Beijing = Peking), Zhejiang (Ningbo = Ning-Po);
Korea :
Kyonggi-do (Kwangnung), Kangwon Do (Mt. Sulak-san).
Bibliography.
ARAYA
K. (2000)
An account of a visit to European Museums. 2. The lucanid specimens in the van
Roon and Siebold collections deposited in the Rijksmuseum van Natuurlijke
Historie, Leiden. Gekkan-Mushi, 350:4-16.
Lucanus
(Lucanus) maculifemoratus ssp. maculifemoratus MOTSCHULSKY, 1861
[Platycerus] [maclifemoratus].
Lucanus
cantori MOTSCHULSKY (nec HOPE) 1860.
Lucanus
sericans VOLLENHOVEN, 1861 [Java, error; in reality Japon, col. Von Siebold].
Lucanus
hopei PARRY, 1862 [Ind. Or., error].
Lucanus
elegans PLANET, 1898.
Lucanus
montivagus MOLLENKAMP, 1912 [Borneo, error].
Lucanus
maculifemoratus f. nakanei KUROSAWA, 1976.
Lucanus
maculifemoratus ssp. maculifemoratus MOTSCHULSKY; Araya, 2000:6-7,
10-11.
Lucanus
sericans VOLLENHOVEN; Araya, 2001:20, figs.73-74.
Lucanus
maculifemoratus maculifemoratus MOTSCHULSKY; Araya, 2001:272.
Note : hopei
= sericans, = elegans is perhaps a valid ssp. for the
specimens of Hokkaido and perhaps North of Honshu.
·
Russia : Kuriles Is. : Kunashir I. = Kunashirito I.
·
Japan, col. Von Siebold, “Muller, Java” (male HOLOTYPUS of Lucanus
sericans VOLLENHOVEN, 1861 in Rijksmuseum, Leiden).
·
Japan : Hokkaido I. :
Ishikari Pref. : Kotoni, 45°05N-141°12E
·
Japan : Hokkaido I. :
Sorachi Pref. : Akabira
·
Japan : Hokkaido I. :
Hidaka : Hidaka
·
Japan : Hokkaido I. :
Oshima Pref. : Hakodate, 41°47N-140°43E
·
Japan : Hokkaido I. :
Gamushi
·
Japan : Hokkaido I. :
Nukabira
·
Japan : Honshu I. :
Akita Pref. : Tobishima I., 39°13N-139°32E
·
Japan : Honshu I. :
Niigata Pref. : Sadogashima I.
·
Japan : Honshu I. : Fukui Pref. (citated
by Araya, 2001).
·
Japan : Honshu I. : Nagano Pref.
·
Japan : Honshu I. : Gifu Pref. : Gifu
·
Japan : Honshu I. : Gifu Pref. : Hirugano
·
Japan : Honshu I. : Gifu Pref. : Mt. Isetera
·
Japan : Honshu I. : Gifu Pref. : Mt. Odai
·
Japan : Honshu I. :
Kanagawa Pref. : Mutsu-ura-cho, 35°20N-139°36E
·
Japan : Honshu I. :
Kanagawa Pref. : Juniso, 35°20N-139°35E
·
Japan : Honshu I. :
Kanagawa Pref. : Sakurayama, 35°17N-139°35E
·
Japan : Honshu I. : Kanagawa
Pref. : Jinmuji, 35°18N-139°36E
·
Japan : Honshu I. :
Kanagawa Pref. : Futago-yama, 35°17N-139°36E
·
Japan : Honshu I. :
Kanagawa Pref. : Takatori-yama, 35°18N-139°36E
·
Japan : Honshu I. :
Kanagawa Pref. : Kumura, 35°14N-139°42E
·
Japan : Honshu I. :
Kanagawa Pref. : Yokohama, 35°20N-139°36E
·
Japan : Honshu I. :
Kanagawa Pref. : Ohgusu-yama, 35°16N-139°37E
·
Japan : Honshu I. :
Kanagawa Pref. : Miura-fuji, 35°12N-139°40E
·
Japan : Honshu I. :
Osaka Pref. : Kobe, 34°42N-135°08E
·
Japan : Honshu I. : Osaka Pref. : Nose : Mt. Myôken
·
Japan : Honshu I. :
Osaka Pref. : Osaka, 34°39N-135°27E
·
Japan : Honshu I. : Tottori Pref. : Mt. Daisen, 35°23N-133°32E : Hooki
·
Japan : Honshu I. : Tottori Pref. : Oki I., 36°14N-133°16E
·
Japan : Honshu I. : Hiroshima Pref. : Tojio cho
·
Japan : Shikoku I. : Ehime Pref. : Matsuyama, 33°50N-132°44E
·
Japan : Shikoku I. : Ehime Pref. : Kanbayashi
·
Japan : Kyushu I. :
Nagasaki Pref. : Tsushima I., 34°20N-129°20E
·
Japan : Kyushu I. : Nagasaki Pref. : Goto Is. : Fukuejima I.,
32°41N-128°44E
·
Japan : Kumage Is. : Kuroshima I.
·
Japan : Kubo
·
Japan : Nooe-Myoken
·
Japan : Clüba
|
|
|
Lucanus maculifemoratus
ssp. maculifemoratus : HOLOTYPUS de Lucanus sericans VOLLENHOVEN
(adapted from Araya, 2000) |
Bibliography.
ARAYA
K. (2000)
An account of a visit to European Museums. 2. The lucanid specimens in the van
Roon and Siebold collections deposited in the Rijksmuseum van Natuurlijke
Historie, Leiden. Gekkan-Mushi, 350:4-16.
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
ARAYA
K. (2001)
New records of Ceruchus lignarius monticola Nakane, 1978 (Coleoptera,
Lucanidae) from Fukui Prefecture, Central Japan, with reference to Lucanid
fauna of Fukui Prefecture. Sukunahikona, Special Publication of the Japan
Coleopterological Society, 1:271-273.
Lucanus
(Lucanus) maculifemoratus ssp. adachii TSUKAWAKI, 1995.
Japan :
Tokyo Met. [Izu Is. (Ooshima I.--34°46N-139°24E {Mt. Miharayama*, Habu*},
Toshima I., Niijima I.--34°24N-139°17E {Mt. Miyazukayama*}, Kozushima I. {Mt.
Tenjosan*, Anakiyama*}, Miyakejima I.--34°04N-139°30E)].
Lucanus
(Lucanus) maculifemoratus ssp. balachowskyi LACROIX, 1968.
Lucanus
balachowskyi LACROIX, 1968 [Japan : Mukoda].
Lucanus
balachowskyi LACROIX; Araya, 2000:14.
Japan : Kyushu I. : Kagoshima Pref. :
Satzuma : Mukoda I.*
Japan :
Prov. Satsuma : Mukoda (1 male TOPOTYPUS of Lucanus balachowskyi LACROIX,
1968 in Rijksmuseum, Leiden).
|
|
|
Lucanus maculifemoratus
ssp. balachowskyi : Mukoda Is. (adapted from Araya, 2000) |
Bibliography.
ARAYA
K. (2000)
An account of a visit to European Museums. 2. The lucanid specimens in the van Roon
and Siebold collections deposited in the Rijksmuseum van Natuurlijke Historie,
Leiden. Gekkan-Mushi, 350:4-16.
Lucanus
(Lucanus) maculifemoratus ssp. ferriei PLANET, 1898 [ferrieri].
Japan : Kagoshima Pref. [Amami-oshima I. (Mt.
Yuwan-dake, Kamiya, Kinsakubaru)].
Lucanus
(Lucanus) maculifemoratus ssp. taiwanus MIWA, 1936.
= Lucanus
ferriei MIWA (nec PLANET) 1931 [ferrieri].
Taiwan* :
Taichung Distr. (Lisan), Hualien Distr.
(Pilu), Nantou Distr. (Sungkang, Wusha, Puli), ? (Kyuhabon, Naihompo, Tompo, Musha).
Lucanus (Lucanus) manai BOMAN & MIYASHITA, 1997.
Lucanus manai BOMAN & MIYASHITA, 1997:2, figs. 5-6.
Boucher,
1998:180.
Myanmar
:
Kachin
Prov., VIII-96, leg. P. Sukkit (male HOLOTYPUS of Lucanus manai BOMAN
& MIYASHITA, female ALLOTYPUS in coll. Miyashita, Japan) (3 males
PARATYPUS, 1 female PARATYPUS in coll. Miyashita, Japan; Bomans, in Natural
History Museum, London; Arnaud, Francia)
|
|
|
Lucanus manai (adapted from
Bomans & Miyashita). |
Bibliography :
BOMAN H. & MIYASHITA T. (1997)
Description de trois nouvelles espèces de Lucanidae de nord Birmanie. 91 ème
contribution à l'étude des Coléoptères Lucanides. Besoiro, 4:2-3.
Lucanus (Lucanus) masumotoi HIRASAWA & AKIYAMA, 1990.
Hirasawa & Akiyama, 1990:55; Imanishi,
1990:15-18; Sakaino & Yu, 1993:15-16; Sakaino, Yu & Chu, 1994:122;
Sakaino, 1994:18-19; Mizunuma & Nagai, 1994:214, pl.12; Sakaino, 1995:3;
Araya, 1996:22.
Lucanus (Lucanus) masumotoi ssp.
masumotoi HIRASAWA &
AKIYAMA, 1990.
Central Taiwan.
Lucanus (Lucanus) masumotoi ssp.
ogakii IMANISHI, 1990.
Lucanus masumotoi ssp. chuyunshanus SAKAINO & YU 1993 [chuyunsanus] [Mt.
Chuyunshan].
Taiwan : Hualien Distr. (Pilu = Piru*,
Taihorin = Tayurin*), Kaohsiung Distr. (Mt. Chuyunshan = Mt. Izumoyama, Mt. Hsi-nan-shan).
Lucanus
(Lucanus) mearesii HOPE, 1842.
Lucanus
mearesii HOPE, 1842:83 [Himalaya].
Lucanus
mearesii HOPE; Hope, 1843:364
Lucanus
mearesii HOPE; Hope, 1845:73
Lucanus
nigripes HOPE & WESTWOOD, 1845:10 [Assam].
Lucanus
mearesii HOPE; Burmeister, 1847:527
Lucanus
mearesii HOPE; Westwood, 1848:21, pl.10
Lucanus
mearesii HOPE; Parry, 1864:72
Lucanus
mearesii HOPE; Planet, 1898:214
Lucanus
mearesii HOPE; Planet, 1902:17, pl.4
Lucanus
mearesii HOPE; Heyne & Taschenberg, 1908:50
Lucanus
mearesii HOPE; Boileau, 1913:219
Lucanus
mearesii HOPE; Stebbing, 1914:72
Lucanus
mearesii HOPE; Gravely, 1915:414
Lucanus
mearesii HOPE; Didier, 1949:pl.19
Lucanus
mearesii HOPE; Arrow, 1950:52, pl.3
Lucanus
mearesii HOPE; Benesh, 1960:145
Lucanus
mearesii HOPE; Kuyten, 1964:141
Lucanus
mearesii HOPE; Kurosawa, 1978:144
Lucanus
mearesii HOPE; Klausnitzer, 1982:42
Lucanus
mearesii HOPE; Sakaino, 1987:16
Lucanus
mearesii HOPE; Maes, 1992:19
Lucanus
mearesii HOPE; Mizunuma & Nagai, 1994:213, pl.10.
Lucanus
mearesii HOPE; Araya, 2001:18, figs.63-64.
·
India : Punjab
·
India : Uttar Pradesh :
Mussoorie
·
India : Sikkim
·
India : West Bengal : Darjeeling
·
India : West Bengal : Kurseong
·
India : West Bengal : Mangpu
·
India : West Bengal : Gopaldhara
·
India : West Bengal : Rungbong Valley
·
India : West Bengal : Ghoom
·
India : Assam (female TYPUS of Lucanus nigripes HOPE &
WESTWOOD, 1845, # 354 in Oxford University Museum of Natural History) [Araya,
2001 wrote that the type comes from ? Tonkin].
·
Himalaya (male TYPUS of Lucanus mearesii HOPE, 1842, # 355 in
Oxford University Museum of Natural History).
·
Nepal
·
Bhutan : Pedong
·
Bangladesh : Sylhet*
Saproxylophagous
: Symplocaceae : Symplocos.
|
|
|
|
male TYPUS of Lucanus mearesii HOPE,
1842, from Himalaya (adapted from Araya, 2001). |
female TYPUS of Lucanus nigripes HOPE
& WESTWOOD, 1845, from Assam (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Lucanus) miwai KUROSAWA, 1966.
Kurosawa,
1966:342; Kurosawa, 1978:143; Kawano, 1988:26; Hashimoto, 1990:23-26, pl.4;
Imanishi, 1990:15; Maes, 1992:19; Sakaino, Yu & Chu, 1994:122; Mizunuma
& Nagai, 1994:217, pl.17; Boucher, 1998:180.
Taiwan :
Nantou Distr. (Sungkang--Tattaka*, Sungkang, Wushe), ? (Mt. Ali-Shan*).
Lucanus miyashitai MIZUNUMA, 1994.
Lucanus
miyashitai MIZUNUMA in MIZUNUMA & NAGAI, 1994:7-8, fig. 3a-c, lam. 17, fig.
119(1-5) [Nan].
Thailand
: Chiang Mai : Nan (male HOLOTYPUS in Kanagawa Prefectural Museum, Japan) (8
male Paratypus in col. Tetsuo Mizunuma, Japan) (2 male Paratypus in col. T.
Miyashita, Japan) (1 male Paratypus in col. H Fujita, Japan) (2 male Paratypus
in col. M. Suzumura, Japan).
Bibliography.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Lucanus (Lucanus) nobilis DIDIER, 1925.
Didier,
1925:221; Didier, 1926:161; Didier & Séguy, 1953:83; Benesh, 1960:145;
Maes, 1992:19; Mizunuma & Nagai, 1994:213-214, pl.11, 138; Boucher,
1998:184.
Vietnam :
Tonkin (Lao-Kay*, Sapa--Hoang Lien So'n).
Lucanus
(Lucanus) parryi BOILEAU, 1899.
Lucanus
parryi BOILEAU, 1899:111
Lucanus
parryi BOILEAU; Planet, 1900:285
Lucanus parryi BOILEAU; Planet, 1902:78
Lucanus parryi BOILEAU; Kriesche, 1922:118
Lucanus parryi ab. aterrimus DIDIER, 1929:91, 95-96, pl.1 [Fokien].
Lucanus parryi ab. thoracicus DIDIER, 1929:91, 95-96, pl.1 [Fokien].
Lucanus laevigatus DIDIER, 1931:225 [Fokien].
Lucanus parryi BOILEAU; Didier, 1949:pl.20
Lucanus parryi BOILEAU; Didier & Séguy, 1953:85
Lucanus parryi BOILEAU; Benesh, 1960:145
Lucanus parryi BOILEAU; Kurosawa, 1978:144-146
Lucanus
parryi BOILEAU; Klausnitzer, 1982:43
Lucanus
parryi BOILEAU; Bomans, 1989:4
Lucanus
parryi BOILEAU; Maes, 1992:19
Lucanus
parryi BOILEAU; Mizunuma & Nagai, 1994:217, pl.17
Lucanus parryi BOILEAU; Kawano, 1997:25.
Lucanus laevigatus DIDIER; Araya, 2001:17, figs. 55-56.
·
China : Yunnan
·
China : Jiangxi : Mt. Zhenzhu Shan
·
China : Zhejiang = Chekiang : Mokanshan
·
China : Zhejiang :
Pamasishi
·
China : Fujian =
Fokien, col. Janson, VI-1906, ex. col. H. Boileau (female TYPUS of Lucanus
laevigatus DIDIER, 1931 in Museum National d’Histoire Naturelle, Paris).
·
China : Fujian = Fokien (TYPUS of Lucanus parryi ab. aterrimus DIDIER,
1929).
·
China : Fujian = Fokien (TYPUS of Lucanus parryi ab. thoracicus DIDIER,
1929).
·
China : Fujian : Hsikong
·
China : Fujian : Atschong
·
China : Fujian : Yungling
·
China : Fujian : Kuatun = Koua-Toun (TYPUS of Lucanus parryi BOILEAU,
1899).
|
|
|
female TYPUS of Lucanus laevigatus DIDIER,
1931, from China : Fujian = Fokien (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus (Lucanus) placidus SAY, 1825 [Pseudolucanus] [America borealis].
Lucanus rupicapra DEJEAN 1837 [nomen nudum] [America borealis].
Lucanus lentus CASTELNAU 1840 [Pseudolucanus] [America
borealis].
Say, 1825:202; Dejean, 1837:193; Castelnau,
1840:171; Burmeister, 1847:356; Leconte, 1859:302; Horn, 1875:137; Fuchs,
1882:51, pl.; Wickham, 1899:22; Meek, 1901:375; Planet, 1902:101; Hart,
1907:244; Heyne & Taschenberg, 1908:50; Blatchley, 1910:906; Girault,
1913:195; Benesh, 1932:40; Milne, 1933:106; Didier & Séguy, 1952:pl.14;
Didier & Séguy, 1953:27; Séguy, 1955:32; Benesh, 1960:145; Ritcher, 1966;
Mathieu, 1969:1055; Blackwelder & Arnett, 1974; Boucher & Huang,
1991:38; Ratcliffe, 1991:267; Maes, 1992:19; Bartolozzi & Onore, 1993:126;
Mizunuma & Nagai, 1994:212, pl.7; Poole & Gentili, 1996:297.
Canada : Ontario (Essex Co.-Pointe
Pelee--41°55N-82°29W, Dunville--42°52N-79°39W, Toronto--43°42N-79°24W);
USA : Oregon, New Mexico, Nebraska [Brown
Co. (Long
Pine--4230N-99°40W), Holt Co. (Spencer Dam--42°50N-98°45W), Franklin
Co.--40°10N-98°45W, Howard Co. (St. Paul--41°15N-98°25W), Madison
Co.--41°45N-97°30W (Meadow Grove), Saline Co. (Crete--40°35N-96°50W), Douglas
Co.--41°15N-95°55W (Waterloo), Cass Co.--41°N-95°50W (Cedar Creek), Cherry Co.
(Dewey Lake), Nuckolls Co., Richardson Co. (Indian Cave State Park), Bellevue],
Kansas, Oklahoma, Minnesota, Iowa, Missouri [Columbia--38°55N-92°20W],
Arkansas, Michigan [Ann Harbor--42°20N-83°45W], Wisconsin [Warck Co.], Illinois
[Chicago--41°50N-87°40W], Indiana [Miller], Kentucky, Mississippi, Alabama,
Ohio, New York [Buffalo--42°55N-78°50W], Pennsylvania [Cornwells], Virginia
[Blacksburg--37°15N-80°30W].
Saproxylophagous : Fagaceae : Quercus.
Lucanus (Lucanus) planeti PLANET, 1899.
Planet, 1899:72; Planet, 1902:49, pl.11;
Didier, 1926:157; Didier, 1949:pl.16; Didier & Séguy, 1953:83; Benesh,
1960:146; De Lisle, 1973:137; Kurosawa, 1978:143; Sakaino, 1981:23;
Klausnitzer, 1982:42; Maes, 1992:19; Bomans, 1994:494; Mizunuma & Nagai,
1994:215, pl.14; Kawano, 1997:454; Kawano, 1997:25.
China :
Sichuan, Yunnan (Kouang-Si-Hien, Yunnan-Sen, Pe-Yen-Tsing, Tse-Tsong);
Vietnam :
Tonkin (Dong-Van*, Tam Dao near Vinh Yen).
Lucanus
(Lucanus) pseudosingularis DIDIER & SEGUY, 1952 (new name for singularis).
Lucanus
singularis DIDIER (nec PLANET) 1949:pl.13.
Lucanus
pseudosingularis DIDIER & SEGUY, 1952:31 (new name for singularis).
Benesh,
1960:146; Lacroix, 1971:553, 562; Lacroix, 1982:16; Maes, 1992:19; Mizunuma
& Nagai, 1994:213.
Lucanus
pseudosingularis DIDIER & SEGUY; Zilioli, 2002:213.
China :
Yunnan* (Kouang-Si-Hien, Mong-Tze, Pe-Yen-Tsing*).
Bibliography
:
ZILIOLI
M. (2002)
Lucanus victorius n. sp., a new outstanding stag beetle from Sichuan,
China (Coleoptera, Lucanidae). Atti Soc. It. Sci. nat. Museo civ. Stor. Nat.
Milano, 143(II):209-213.
Lucanus
(Lucanus) pulchellus DIDIER, 1925.
Lucanus
pulchellus DIDIER, 1925:220 [Vietnam : Tonkin].
Lucanus
pulchellus DIDIER; Didier, 1926:157
Lucanus
pulchellus DIDIER; Didier & Séguy, 1953:85
Lucanus
pulchellus DIDIER; Benesh, 1960:146
Lucanus
pulchellus DIDIER; Maes, 1992:19
Lucanus
pulchellus DIDIER; Mizunuma & Nagai, 1994:216-217, pl.16, 138.
Lucanus
pulchellus DIDIER; Araya, 2001:19, figs.66-70.
·
Vietnam : Tonkin (male TYPE, female TYPE of Lucanus pulchellus DIDIER,
1925 in Museum National d’Histoire Naturelle, Paris).
·
Vietnam : Tonkin : Sapa : Hoang Lien So'n.
|
|
|
|
male TYPE of Lucanus pulchellus DIDIER,
1925, from Tonkin (adapted from Araya, 2001). |
female TYPE of Lucanus pulchellus DIDIER,
1925, from Tonkin (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Lucanus) datunensis HASHIMOTO, 1984.
Lucanus
datunensis HASHIMOTO, 1984:18 [Taiwan : Mt. Datun].
Lucanus
ritae LACROIX, 1984:10 [Mt. Ali-shan].
Lucanus
ritae LACROIX; Lacroix, 1984:5.
Lucanus datunensis HASHIMOTO; Sakaino, 1985:15-18
Lucanus datunensis HASHIMOTO; Nagai, 1985.
Lucanus datunensis HASHIMOTO; Kawano, 1988:26
Lucanus datunensis HASHIMOTO; Hashimoto, 1990:23-26, pl.4
Lucanus datunensis HASHIMOTO; Wang, 1990.
Lucanus datunensis HASHIMOTO; Tanikado, 1992.
Lucanus datunensis HASHIMOTO; Chang, 1993.
Lucanus datunensis HASHIMOTO; Sakaino & Yu, 1994:26-27
Lucanus datunensis HASHIMOTO; Sakaino, Yu & Chu, 1994:122
Lucanus datunensis HASHIMOTO; Mizunuma & Nagai, 1994:217, pl.16.
Lucanus datunensis HASHIMOTO; Wang, 1994.
Lucanus datunensis HASHIMOTO; Araya, 2000:1-4.
Lucanus ritae LACROIX; Araya, 2001:19-20, fig.71.
Lucanus datunensis HASHIMOTO; Araya, 2001:19.
·
Taiwan : Taipei Distr. : Mt. Tatun Shan = Mt. Datun (male HOLOTYPUS of Lucanus
datunensis HASHIMOTO, 1984).
·
Taiwan : Central Taiwan : Mt. Ali Shan (male HOLOTYPUS of Lucanus
ritae LACROIX, 1984) (2 males PARATYPUS of Lucanus ritae LACROIX,
1984 in Museum d’Histoire Naturelle de Geneve).
|
|
|
|
Lucanus datunensis : male HOLOTYPUS
(adapted from Araya) |
|
|
|
|
|
Lucanus datunensis : male HOLOTYPUS
of Lucanus ritae (adapted from Araya) |
Lucanus datunensis : male PARATYPUS
of Lucanus ritae (adapted from Araya) |
Bibliography.
ARAYA
K. (2000)
Notes on Lucanus ritae Lacroix, 1984, a junior synonym of Lucanus
datunensis Hashimoto, 1984. Kogane, Tokyo, 1:1-4.
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
CHANG
Y. (1993)
Stag beetles of Taiwan. Newton Press, Taipei, 111 pp. [in Chinese].
HASHIMOTO
S. (1984)
A new species of the genus Lucanus of the family Lucanidae inhabiting
Taiwan. Iwase, 2:23-26.
LACROIX J.P. (1984) Descriptions de Coleoptera Lucanidae nouveaux
ou peu connus. Bull. Sciences Nat, 40(1983):5-18, 7 pls.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanidae of the world. Mushi-sha’s
Iconographic Series of Insects. 1:337 pp. Mushi-sha, Tokyo.
NAGAI
S. (1985)
Coleopterorum Catalogus, World Lucanidae. Mokuyo-sha, Tokyo. 224 pp. [In
Japanese].
SAKAINO
H. (1985)
Miscellaneous notes on a Formosan Lucanidae, Lucanus datunensis Hasimoto.
Gekkan-Mushi, 172:15-18.
SAKAINO
H. & YU C.K. (1994) The female of Lucanus datunensis Hashimoto
(Coleoptera, Lucanidae). Gekkan-Mushi, 285:26-27.
TANIKADO
M. (1992)
Current topics of stag beetles in Taiwan. Gekkan-Mushi, 256:22-25.
WANG
H.Y. (1990) Illustrations of stag beetles in Taiwan. Taiwan Museum, Taipei,
189 pp. (In Chinese).
WANG
H.Y. (1994) Illustrations of stag beetles in Taiwan. Taiwan Museum, Taipei,
207 pp. (In Chinese).
Lucanus
(Lucanus) sericeus DIDIER, 1925.
Lucanus
sericeus DIDIER, 1925:219 [Tonkin : Dong-Dang].
Lucanus sericeus DIDIER; Mizunuma & Nagai, 1994:213, pl.11
Lucanus sericeus DIDIER; Kawano, 1997:454
Lucanus sericeus DIDIER; Kawano, 1997:25.
Lucanus sericeus DIDIER; Araya, 2001:20-21, figs.75-78.
·
Thailand : Chiang Mai : Fang : Mt. Doi Pahompok
·
Vietnam : Tonkin : Dong-Dang, V-1915, col. R. Vitalis de Salvaza, ex. col. H. Boileau, ex. col. R. Didier (male
TYPE of Lucanus sericeus DIDIER, 1925 in Museum National d’Histoire
Naturelle, Paris).
·
Vietnam : Tonkin : Chapa, VI-1915, col. R. Vitalis de Salvaza, ex. col. R. Didier (female COTYPE of Lucanus
sericeus DIDIER, 1925 in Museum National d’Histoire Naturelle, Paris).
|
|
|
|
male TYPE of Lucanus sericeus DIDIER,
1925, from Tonkin (adapted from Araya, 2001). |
female COTYPE of Lucanus sericeus DIDIER,
1925, from Tonkin (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Lucanus) singularis PLANET, 1899 [Indes Orientales].
Lucanus
furcifer ARROW 1950 [Sikkim--Lachen Lochung, Yunnan, Szechuan--Ouy-Sy].
Planet,
1899; Planet, 1900:11; Planet, 1902:22; Planet, 1903:12; Didier, 1949:pl.13;
Arrow, 1950:46, 62, pl.3; Didier & Séguy, 1953:29-30; Benesh, 1960:141,
146; Lacroix, 1971:553, 566; De Lisle, 1975; Maes, 1992:19; Mizunuma &
Nagai, 1994:213.
India :
Sikkim (Lachen Lachung);
Bhutan :
Tongsa;
China :
Sichuan (Ouy-Sy), Xizang = Thibet (Zayul), Yunnan (Mit-Sao).
Lucanus
(Lucanus) smithii PARRY, 1862 [smithi].
Lucanus
vicinus PLANET (nec HOPE & WESTWOOD) 1900.
Parry,
1862:108; Parry, 1864:10, pl.10; Planet, 1900:164; Planet, 1902:63, pl.14;
Heyne & Taschenberg, 1908:50; Boileau, 1913:219; Gravely, 1915:413; Arrow,
1950:49, pl.4-5; Didier & Séguy, 1952:pl.20; Benesh, 1960:146; Kurosawa,
1970:160; Lacroix & Bomans, 1973:145; Maes, 1992:19; Bartolozzi &
Sforzi, 1994:114; Mizunuma & Nagai, 1994:214, 309, pl.12.
India* :
Sikkim (Tendong), West Bengal (Darjeeling, Kurseong, Mangpu, Gopaldhara, Rungbong
Valley, Siliguri, Terai), Assam, Maharashtra (Punah);
China :
Yunnan (Mit-Sao);
? Nepal;
Bhutan :
Pedong, Sampa-Kotoka;
Bangladesh
: Sylhet.
Lucanus (Lucanus) speciosus DIDIER, 1925.
Lucanus speciosus DIDIER, 1925:220 [Tonkin:Chapa].
Lucanus speciosus DIDIER; Didier, 1926:157
Lucanus speciosus DIDIER; Didier & Séguy, 1953:87
Lucanus speciosus DIDIER; Benesh, 1960:146
Lucanus speciosus DIDIER; Kurosawa, 1978:143
Lucanus speciosus DIDIER; Maes, 1992:20.
Lucanus speciosus DIDIER; Araya, 2001:21, figs.79-83.
·
Vietnam : Tonkin : Chapa, VI-1916 (male TYPUS, female COTYPUS of Lucanus
speciosus DIDIER, 1925 in Museum National d’Histoire Naturelle, Paris).
|
|
|
|
male TYPUS of Lucanus speciosus DIDIER,
1925, from Tonkin (adapted from Araya, 2001). |
female COTYPUS of Lucanus speciosus DIDIER,
1925, from Tonkin (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae) from
Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Lucanus) swinhoei PARRY, 1874.
Parry,
1874:370, pl.4; Planet, 1902:88, pl.15; Miwa, 1931:321; Benesh, 1960:146; Kurosawa,
1978:143; Hashimoto, 1990:23-26, pl.4; Maes, 1992:20; Sakaino, Yu & Chu,
1994:122; Mizunuma & Nagai, 1994:216, pl.16; Araya, 1996:22.
Taiwan = Formosa* : Taipei Distr. (Mt. Tatun Shan),
Taichung Distr. (Lisan), Nantou Distr. (Sungkang, Yushin,
Puli), ? (Sinpeitou).
Lucanus
(Lucanus) szetschuanicus HANUS, 1932.
Lucanus
szetschuanicus HANUS, 1932:101 [China : Sichuan = Szetschuan].
Lucanus
szetschuanicus HANUS; Hanus, 1935:23
Lucanus
szetschuanicus HANUS; Didier & Séguy, 1952:pl.19
Lucanus
szetschuanicus HANUS; Benesh, 1960:146
Lucanus
szetschuanicus HANUS; Maes, 1992:20
Lucanus
szetschuanicus HANUS; Mizunuma & Nagai, 1994:216, pl.16.
Lucanus
szetschuanicus HANUS; Araya, 2001:21, figs.84-85.
·
China : Sichuan = Szetschuan : Giufu-Schan, leg. E. Reitter (female
PARATYPUS of Lucanus szetschuanicus HANUS, 1932 in Museum National
d’Histoire Naturelle, Paris).
·
China : Sichuan =
Szetschuan : Leshan
|
|
|
female PARATYPUS of Lucanus szetschuanicus HANUS,
1932, from Szetschuan (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.
Lucanus
(Lucanus) tetraodon THUNBERG, 1806.
Thunberg,
1806:188, 197; Schonherr, 1817:232; Dejean, 1837:193; Burmeister, 1847:349;
Costa, 1853:17; Duval, 1859:11; Fairmaire, 1859:275; Gautier des Cottes,
1860:53; Kraatz, 1860:68, 265-266, pl.7; Parry, 1864:72; Motschulsky, 1870:42,
44; Reitter, 1891:144; Planet, 1899:89, 98, pl.16; Planet, 1902:126; Barthe,
1909:5; Leoni, 1910:155; Schaufuss, 1916:1253; Portevin, 1931:3; Porta,
1932:371; Colas, 1949:128-131; Didier, 1949:pl.6; Bourgin, 1950:66; Didier
& Séguy, 1953:87; Benesh, 1960:147; Colas, 1962:118; Colas, 1970:77;
Lacroix & Bomans, 1973:148; Paulian, 1978:7; Bartolozzi, 1986:17; Paulian,
1986:91; Bartolozzi, 1989:104; Maes, 1992:20; Bartolozzi & Onore, 1993:126;
Mizunuma & Nagai, 1994:212, pl.7.
Lucanus (Lucanus) tetraodon ssp.
provincialis COLAS, 1949.
S. France : Var (Seyne-sur-Mer--43°05N-5°53E--Forêt
de Janas*).
Lucanus (Lucanus) tetraodon ssp.
corsicus GAUTIER DES COTTES,
1860.
France : Corse* (Mont d'Oro, Bastia, Murato,
Rutali, Pont de Castirla near Francardo, San Nicolo-di-Moriani, Eaux d'Orezza
near Piedicrocce, Cuttoli-Corticchiato near Ajaccio, Bonifacio, Bastia, Lento
at 45 km S of Bastia, Tavera, Calvi, Cuttoli-Corticchiato at N of Ajaccio, Col
de San Sebastiano, Casabianda near Aléria, Zicavo).
Lucanus
(Lucanus) tetraodon ssp. tetraodon THUNBERG, 1806 [Platycerus].
Lucanus
bidens THUNBERG, 1806 [Italia].
Lucanus impressus THUNBERG, 1806 [Italia].
Lucanus barbarossa (FABRICIUS); COSTA, 1853 [Italia].
Lucanus
serraticornis FAIRMAIRE, 1859 [Italia].
Lucanus
tetraodon THUNBERG; Bartolozzi, 1986.
Lucanus
tetraodon THUNBERG; Carpaneto & Piatella, 1995.
Lucanus
tetraodon THUNBERG; Franciscolo, 1997.
Lucanus
tetraodon THUNBERG; Carpaneto, Maltzeff, Piatella & Pontuale, 1998:16-17.
Lucanus
tetraodon THUNBERG; Carpaneto, Maltzeff, Piatella & Facchinelli, 2001:312,
326.
Lucanus
tetraodon THUNBERG; Ballerio, 2003:68, 73, 82.
Lucanus
tetraodon THUNBERG; Zilioli & Pittino, 2004:301-306, figs. 1-3.
Ø
N. Italy : Lombardia :
Milano : Ticino River : Turbigo, 10-VII-1953, leg. G. Mariani (1 male in col. Riccardo Pittino, Italia).
Ø
N. Italy : Lombardia : Varese : Gallerate dintorni ovest : Castelnovate,
17-VII-2000, leg. L. & R. Pittino (1 male in col. Riccardo Pittino, Italia).
Ø
N. Italy : Lombardia :
Varese : Gallerate dintorni ovest : Castelnovate, 12-VIII-2004, leg. P. Gandini
(2 males in col. Michele Zilioli, Italia).
Ø
N. Italy : Lombardia :
Varese : Gallerate dintorni ovest : Vizzola Ticino, 200 m, 25-VII-2001, leg. R. Pittino (2
males in col. Riccardo Pittino, Italia).
Ø
N. Italy : Lombardia :
Varese : Gallerate dintorni ovest : Vizzola Ticino, 200 m, 25-VII-2001, leg. M. Zilioli (4
males in col. Michele Zilioli, Italia).
Ø
N. Italy : Lombardia :
Varese : Gallerate dintorni ovest : Vizzola Ticino, 200 m, 30-VII /
3-VIII-2001, leg. M. Zilioli & P. Gandini (15 males in col. Michele Zilioli, Italia).
Ø
N. Italy : Lombardia :
Varese : Gallerate dintorni ovest : Vizzola Ticino, 200 m, 6-VIII-2004, leg. M. Zilioli (7
males, 1 female in col. Michele Zilioli,
Italia).
Ø N. Italy : Lombardia : Varese : Gallerate dintorni
ovest : Vizzola Ticino, 200 m, 29-VII-2004, leg. M. Zilioli & P. Gandini
(23 males, 3 females in col. Michele Zilioli, Italia and Museo Civico di Storia
Naturale di Milano).
Ø Italy : Firenze
(citated by Carpaneto & Piatella, 1995; Franciscolo, 1997).
Ø S. Italy (TYPUS de
Lucanus bidens THUNBERG, 1806).
Ø
S. Italy : Umbria :
Terni, 42°34N-12°40E : Piediluco.
Ø
S. Italy : Abruzzes :
Tagliacozzo, 42°05N-13°15E.
Ø
S. Italy : Basilicata :
Potenza, 40°39N-15°47E : Picena
Ø
S. Italy : Roma :
Castelporziano, 8-XII-67, leg. E. Migliaccio (in Carpaneto, Maltzeff, Piatella
& Pontuale, 1998)
Ø
S. Italy : Roma :
Castelporziano, VI-87, leg. E. Pacieri (in coll. Tenuta Presidenziale, Italia).
Ø
S. Italy : Roma :
Castelporziano, 6-VIII-97, leg. L. Salvati (in coll. L. Salvati, Roma, Italia).
Ø S. Italy : Roma : Castelfusano, 13-VII-2001, leg. L.
Facchinelli (in Carpaneto, Maltzeff, Piattella & Facchinelli, 2001).
Ø S. Italy : Roma :
Capocotta (in Carpaneto, Maltzeff, Piatella & Pontuale, 1998).
Ø Albania.
Ø Greece.
Note :
this species is protected by Regional Law of Toscana, Italia, appendix A and B
(Ballerio, 2003).
Saproxylophagous : Fagaceae : Castanea.
Bibliography :
BALLERIO A. (2003) EntomoLex : la conservazione degli insetti e
la legge. Memorie Soc. entomol. Italiana, 82(1):17-86.
BARTOLOZZI L. (1986) Note
corologiche e morfologiche sui Lucanidae in Toscana (Coleoptera). Atti del
Museo Civico de Storia Naturale di Grosseto 1986(7/8): 11-26.
CARPANETO G.M. &
PIATTELLA E. (1995)
Checklist delle specie della Fauna Italiana. 50. Coleoptera Polyphaga V
(Lucanoidea, Scarabaeoidea). Ed. Calderini, Bologna.
CARPANETO G.M., MALTZEFF
P., PIATELLA E. & FACHINELLI L. (2001) Nuovi reperti di Coleotteri Lamelicorni della Tenuta
Presidenziale di Castelporziano e delle aree limitrofe (Coleoptera,
Lamellicornia). Bollettino dell'Associazione Romana di Entomologia,
56(1-4):311-329.
FRANCISCOLO M.E. (1997) Fauna d’Italia. Vol. XXXV. Coleoptera
Lucanidae. Ed. Calderini, Bologna.
ZILIOLI M. & PITTINO R. (2004) Un reperto eccezionale: Lucanus tetraodon Thunberg
in Lombardia (Coleoptera, Lucanidae). Atti della Societa Italiana di Scienze
Naturali e del Museo Civico di Storia Naturale in Milano, 145(II):301-306.
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Lucanus tetraodon ssp. tetraodon : Italy : Lombardia (adapted from Zilioli and
Pittino). |
Lucanus (Lucanus) tetraodon ssp.
sicilianus PLANET, 1899 [siciliana].
Italy : Sicilia*--37°35N-14°15E.
Lucanus (Lucanus) tetraodon ssp.
argeliensis MAES, 1995.
Lucanus barbarossa BURMEISTER (nec FABRICIUS) 1847.
Algeria.
BIBLIOGRAFIA
CARPANETO G.M., MALTZEFF
P., PIATELLA E. & PONTUALE G. (1998) I Coleotteri Lamellicorni della Tenuta
Presidenziale di Castelporziano e delle aree limitrofe (Coleoptera,
Lamellicornia). Bollettino dell'Associazione Romana di Entomologia,
52(1997):9-54.
Lucanus (Lucanus) thibetanus ssp.
thibetanus PLANET, 1898.
Planet, 1898:277; Planet, 1902:24; Benesh,
1960:147; Lacroix, 1971:553, 563; Maes,
1992:20.
Lucanus tibetanus tibetanus PLANET, Mizunuma & Nagai, 1994:7, fig. 2, 213,
pl. 11, 138
China :
Sichuan, Xizang = Thibet (Manang-Kouang, Siao-Lou* = Sia-Lou Chen), Yunnan.
Bibliography.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Lucanus
(Lucanus) thibetanus ssp. katsurai MIZUNUMA, 1994.
Lucanus
tibetanus ssp. katsurai MIZUNUMA in MIZUNUMA & NAGAI, 1994:7, fig. 1, lam.
138, figs. 92-93 [Sapa].
v N. Vietnam : Hoang
Lien So'n : Sapa (male Holotypus in col. Kanagawa Prefectural Museum, Japan).
Bibliography.
MIZUNUMA
T. & NAGAI S. (1994) The Lucanid Beetles of the World. Mushi-Sha's
Iconographic Series of Insects, Ed. H. Fujita, Japan. Vol. I : 337 pp., 156
pls.
Lucanus
(Lucanus) victorius ZILIOLI, 2002.
Lucanus
(Lucanus) victorius ZILIOLI, 2002:209-213, figs. 1-3.
v China : Sichuan :
Daxue Shan Mts : Yinjing, V/VI-2002, col. K. Fukinuki (male HOLOTYPUS of Lucanus
victorius ZILIOLI 2002 in Museo Civico di Storia Naturale di Milano; 5
males PARATYPUS, 1 female PARATYPUS in col. K. Fukinuki, Hiroshima, Japan; 1
male PARATYPUS in col. N. Okuda, Osaka, Japan; 1 male PARATYPUS in col. M.
Suzumura, Kobe, Japan; 1 male PARATYPUS in col. M. Zilioli, Italy).
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Lucanus victorius : male
HOLOTYPUS, female PARATYPUS (adapted from Zilioli, 2002). |
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Lucanus
victorius : male genitalia. A.dorsal view, b.apex
of flagellum, c.ventral view, d.lateral view (adapted from Zilioli, 2002). |
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Lucanus victorius : male
gonosomite. A.dorsal view, b.ventral view (adapted from Zilioli, 2002). |
Bibliography
:
ZILIOLI
M. (2002)
Lucanus victorius n. sp., a new outstanding stag beetle from Sichuan,
China (Coleoptera, Lucanidae). Atti Soc. It. Sci. nat. Museo civ. Stor. Nat. Milano, 143(II):209-213.
Lucanus (Lucanus) villosus HOPE, 1831.
Hope, 1831:22; Hope & Westwood, 1845:9;
Parry, 1863:446, 453; Planet, 1898:165; Planet, 1902:7; Boileau, 1913:219;
Arrow, 1950:50, pl.4-5; Didider & Séguy, 1953:86; Benesh, 1960:147;
Endrodi, 1968:50; Endrodi, 1971:10-11; Lacroix, 1971:553, 559; Maes, 1992:20;
Bartolozzi & Sforzi, 1994:112, 114; Mizunuma & Nagai, 1994:213, pl.11.
Nepal* : Thodung, Chalsa, Okaldunka, Mt.
Pulchok--Godavari, Bhakta, Lalitpur Distr.--Phulcoki, Sunkhuwasabha
Distr.--Koshi Zone--Chauki, Terhathum Distr.--Koshi Zone--Basantapur, Taplejung
Distr.--Mechi Zone--Gurja;
China :
Yunnan (Yunnan-Sen).
Lucanus
(Lucanus) westermanni HOPE & WESTWOOD, 1845.
Lucanus
westermanni HOPE & WESTWOOD, 1845:10 [Assam].
Lucanus
westermanni HOPE & WESTWOOD; Planet, 1899:34
Pseudolucanus
mniszechi PLANET, 1900:96 [Ind. Or.].
Lucanus
westermanni HOPE & WESTWOOD; Planet, 1902:3, 100, pl.1
Lucanus
westermanni HOPE & WESTWOOD; Heyne & Taschenberg, 1908:50
Lucanus
westermanni HOPE & WESTWOOD; Boileau, 1913:219
Lucanus
westermanni HOPE & WESTWOOD; Gravely, 1915:413
Lucanus
westermanni HOPE & WESTWOOD; Arrow, 1935:106
Lucanus
westermanni HOPE & WESTWOOD; Arrow, 1950:57, pl.4
Lucanus
westermanni HOPE & WESTWOOD; Didier & Séguy, 1952:pl.16
Lucanus
westermanni HOPE & WESTWOOD; Benesh, 1960:148
Lucanus
westermanni HOPE & WESTWOOD; Lacroix & Bomans, 1973:144-145
Lucanus
westermanni HOPE & WESTWOOD; Maes, 1992:20
Lucanus
westermanni HOPE & WESTWOOD; Bartolozzi & Sforzi, 1994:114
Lucanus
westermanni HOPE & WESTWOOD; Mizunuma & Nagai, 1994:214, 309, pl.11.
Lucanus
westermanni HOPE & WESTWOOD; Araya, 2001:21-22, fig.86.
·
India : Sikkim
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India : West Bengal : Darjeeling
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India : West Bengal : Kurseong
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India : West Bengal : Gopaldhara
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India : West Bengal : Rungbong Valley
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India : West Bengal : Mangpu
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India : Assam (male TYPE of Lucanus westermanni HOPE &
WESTWOOD, 1845 # 353 in Oxford University Museum of Natural History) [Araya
wrote “?Himalaya” as type locality].
·
Nepal : Karikhola
·
Nepal : Dudh Kosi-Tal
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Nepal : Junbesi
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Nepal : Kuinibisona
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Bhutan : Pedong
·
China : Sichuan : Siao-Lou
·
China : Fujian : At Chong
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male TYPE of Lucanus westermanni HOPE
& WESTWOOD, 1845, from Assam or Himalaya (adapted from Araya, 2001). |
Bibliography
:
ARAYA
K. (2001)
Notes on type specimens of the genus Lucanus (Coleoptera, Lucanidae)
from Asia stored in several European Museum (1). Gekkan-Mushi, 362:8-22.